Geographic Range
Cygnus atratus
, commonly known as black swans, are native to Australia (including Tasmania) and
have been introduced to New Zealand, Europe, and North America. Black swans are found
mainly in the wetlands of southern Australia and tend to avoid the northern tropics.
They can also be found in across the rest of southern Australia, and in the southeast
of Tasmania. After being introduced to Europe as pets, they can now be found there
in the wild.
- Biogeographic Regions
- nearctic
- palearctic
- australian
- oceanic islands
Habitat
Black swans live in lakes, rivers and swampland, which can be fresh, salt or brackish
water. They prefer habitats with aquatic vegetation. While their natural habitat
is aquatic they are sometimes found in terrestrial areas such as dry pastures or flooded
fields when food is scarce.
- Habitat Regions
- temperate
- tropical
- terrestrial
- Aquatic Biomes
- lakes and ponds
- rivers and streams
- coastal
- brackish water
- Wetlands
- swamp
- Other Habitat Features
- agricultural
- riparian
- estuarine
Physical Description
Swans are the largest of all waterfowl. Black swans' closest relatives are mute swans
(
Cygnus olor
).
Cygnus atratus
has the classical swan look with a long arched neck and raised eyebrows. As the
name implies they are mostly black. Some of the wing feathers are white. They also
have reddish or pinkish irises and richly colored red bills with a white line. The
juveniles are greyish brown with light tipped feathers and a lighter colored bill.
As with many birds, there is sexual dimorphism where the male (called a "cob") is
slightly larger than the female (called a "pen").
When they are fully grown they have a length of 110 to 140 cm and weigh between 3700
to 8750 g. The wingspan ranges between 160 to 200 cm.
- Other Physical Features
- endothermic
- bilateral symmetry
- Sexual Dimorphism
- male larger
Reproduction
Black swans are monogamous and often have the same mate for life. They are territorial
and stay in solitary pairs when mating but are known to occasionally mate in colonies.
The threatening behavior of black swans is similar to mute swans; they both flap and
wave their wings with two or three strokes followed by a call. However, the wings
of black swans make a louder sound than mute swans. Also the standing stance is different;
black swans hold their necks erect with a downward point of the bill and ruffled feathers.
One particularly interesting thing about the courting behavior of black swans is the
"Triumph Ceremony". It is used to strengthen pair-bonds between mates, between parents
and cygnets (baby swans), and for threatening territorial displays. The male swan
approaches the female swan with wings and chin lifted, calling repeatedly. Then the
female returns the same call. They then dip their heads alternating with erect postures.
After this the birds call with their necks outstretched and bills pointed upward;
then they hold their necks at a forty five degree angle and point their bills downward
and at a right angle, they proceed to swim in a circle. These ceremonies are primarily
initated by the male and tend to increase in frequency when there are more swans around.
- Mating System
- monogamous
The breeding season is from February through September. Usually the female (occasionally
the male) makes a nest of sticks, dead leaves and debris into a floating mound on
top of the water. Each female may lay between 5 to 6 eggs, the eggs are laid one
day apart. There is a 35 to 48 day incubation period which begins when all the eggs
have been laid. Males are known to help with incubation. Chicks are precocial but
are brooded on the nest for 2 to 3 weeks after hatching. They fledge from 150 to 170
days after hatching. They remain in family groups for about 9 months and are able
to fly at around 6 months old. The chicks are sexually mature in 18 to 36 months.
Young black swans join juvenile flocks for one to two years before they begin breeding.
- Key Reproductive Features
- iteroparous
- seasonal breeding
- gonochoric/gonochoristic/dioecious (sexes separate)
- sexual
- fertilization
- oviparous
Both male and female black swans incubate the eggs. Chicks are precocial and can
swim and feed soon after hatching. They may ride on their parents' backs when they
venture into deep water. The chicks can fly in 2 months, but they remain in the family
group until the next breeding season. Juvenile black swans often form flocks until
they find a mate.
- Parental Investment
- no parental involvement
- precocial
-
pre-hatching/birth
-
protecting
- male
- female
-
protecting
-
pre-weaning/fledging
-
protecting
- male
- female
-
protecting
-
pre-independence
-
protecting
- male
- female
-
protecting
Lifespan/Longevity
Black swans have been known to live for forty years in the wild.
Behavior
Black swans tend to move in flocks; they are the least territorial of all swans and
sometimes nest in colonies. They are nomadic when food is scarce but are otherwise
sedentary. They feed at dusk and travel at night, calling as they fly, but most activity
is during the day.
- Key Behaviors
- flies
- natatorial
- diurnal
- motile
- nomadic
- sedentary
- territorial
- social
- colonial
Home Range
We do not have information on home range for this species at this time.
Communication and Perception
Black swans use calls and visual signals to communicate. They have advertisment calls
used in territorial defense and specific calls used in Triumph Ceremonies. They have
a high pitched, weak voice. They also use visual displays to communicate such as
raising their shoulders or flapping their wings to threaten predators or other swans
in their territory.
- Other Communication Modes
- duets
Food Habits
Cygnus atratus
eats sub-aquatic foliage that it can reach under water using its long neck. It is
herbivorous, eating vegetation and plants either in the water or on land in pastures
or on farm land. Some common aquatic plants that they feed on are:
Typha
,
Potamogeton
,
Myriophyllum
,
Ruppia
and algae. Occasionally they also eat insects.
- Animal Foods
- insects
- Plant Foods
- leaves
- algae
Predation
Black swans flap their wings which produce loud noises and threaten predators with
their necks erect and bills pointed down. Eggs are taken by
Australian ravens
,
common rats
and
golden-bellied water rats
,
swamp harriers
,
white-bellied sea eagles
, and other hawks. Fledglings are preyed on by swamp harriers, white-bellied sea eagles,
quolls
, golden-bellied water rats, and sometimes gulls and terns.
Ecosystem Roles
Black swans are important members of thier eosystem both as a predator and as prey for other species.
Economic Importance for Humans: Positive
Humans benefit from black swans because they eat their eggs and keep them as pets.
They are also popular among birdwatchers.
- Positive Impacts
- pet trade
- food
- ecotourism
Economic Importance for Humans: Negative
Black swans are common crop pests, either destroying vegetation or uprooting it. In
order to help control black swan populations, a hunting season has been established
in some areas.
- Negative Impacts
- crop pest
Conservation Status
Currently, black swans are not suffering from population declines. Populations range
from the thousands up to tens of thousands in New South Wales.
Additional Links
Contributors
Alaine Camfield (editor), Animal Diversity Web.
Cheryl Jackson (author), University of Michigan-Ann Arbor, Phil Myers (editor), Museum of Zoology, University of Michigan-Ann Arbor.
- Nearctic
-
living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.
- introduced
-
referring to animal species that have been transported to and established populations in regions outside of their natural range, usually through human action.
- Palearctic
-
living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.
- introduced
-
referring to animal species that have been transported to and established populations in regions outside of their natural range, usually through human action.
- Australian
-
Living in Australia, New Zealand, Tasmania, New Guinea and associated islands.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- oceanic islands
-
islands that are not part of continental shelf areas, they are not, and have never been, connected to a continental land mass, most typically these are volcanic islands.
- introduced
-
referring to animal species that have been transported to and established populations in regions outside of their natural range, usually through human action.
- temperate
-
that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).
- tropical
-
the region of the earth that surrounds the equator, from 23.5 degrees north to 23.5 degrees south.
- terrestrial
-
Living on the ground.
- coastal
-
the nearshore aquatic habitats near a coast, or shoreline.
- brackish water
-
areas with salty water, usually in coastal marshes and estuaries.
- swamp
-
a wetland area that may be permanently or intermittently covered in water, often dominated by woody vegetation.
- agricultural
-
living in landscapes dominated by human agriculture.
- riparian
-
Referring to something living or located adjacent to a waterbody (usually, but not always, a river or stream).
- estuarine
-
an area where a freshwater river meets the ocean and tidal influences result in fluctuations in salinity.
- monogamous
-
Having one mate at a time.
- iteroparous
-
offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).
- seasonal breeding
-
breeding is confined to a particular season
- sexual
-
reproduction that includes combining the genetic contribution of two individuals, a male and a female
- fertilization
-
union of egg and spermatozoan
- oviparous
-
reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.
- young precocial
-
young are relatively well-developed when born
- natatorial
-
specialized for swimming
- diurnal
-
- active during the day, 2. lasting for one day.
- motile
-
having the capacity to move from one place to another.
- nomadic
-
generally wanders from place to place, usually within a well-defined range.
- sedentary
-
remains in the same area
- territorial
-
defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement
- social
-
associates with others of its species; forms social groups.
- colonial
-
used loosely to describe any group of organisms living together or in close proximity to each other - for example nesting shorebirds that live in large colonies. More specifically refers to a group of organisms in which members act as specialized subunits (a continuous, modular society) - as in clonal organisms.
- visual
-
uses sight to communicate
- acoustic
-
uses sound to communicate
- duets
-
to jointly display, usually with sounds in a highly coordinated fashion, at the same time as one other individual of the same species, often a mate
- pet trade
-
the business of buying and selling animals for people to keep in their homes as pets.
- food
-
A substance that provides both nutrients and energy to a living thing.
- ecotourism
-
humans benefit economically by promoting tourism that focuses on the appreciation of natural areas or animals. Ecotourism implies that there are existing programs that profit from the appreciation of natural areas or animals.
- herbivore
-
An animal that eats mainly plants or parts of plants.
- folivore
-
an animal that mainly eats leaves.
- endothermic
-
animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.
- bilateral symmetry
-
having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.
- visual
-
uses sight to communicate
- tactile
-
uses touch to communicate
- acoustic
-
uses sound to communicate
- chemical
-
uses smells or other chemicals to communicate
References
Delacour, J. 1954. Waterfowl of the World . London: Country Life Limited.
Forshaw, J. 1998. Aniseriformes. Pp. 84 in Encyclopedia of Birds , Vol. 1, 2nd Edition. McMahons Point, N.S.W.: Weldon Owen.
Johnsgard, P. 1965. Handbook of Waterfowl Behavior . Ithaca, New York: Comstock Publishing Associates.
Kraaijeveld, K., R. Mulder. 2002. The Functions of the Triumph Ceremonies in the Black Swan. Behavior , 139(1): 45-54.
Wilmore, S. 1974. Swans of the World . New York, New York: Taplinger Publishing Co..
del Hoyo, J., A. Elliott, J. Sargatal. 1992. Black Swan. Pp. 578 in Handbook of the Birds of the World , Vol. 1, 1st Edition. Barcelona: Lynx Edicions.
Sedgwick County Zoo. 2003. "Black Swan (Cygnus atratus)" (On-line). Accessed April 06, 2004 at http://www.scz.org/animals/s/bswan.html .
The Chaffee Zoo. Date Unknown. "Black Swan" (On-line). Accessed April 06, 2004 at http://www.chaffeezoo.org/animals/blackSwan.html .