Geographic Range
Himalayan gorals range throughout the Himalayas from Bhutan to Pakistan.
- Biogeographic Regions
- palearctic
- oriental
Habitat
Naemorhedus goral
occupies the forests and scrub-covered slopes of the of the Himalayas. Himalayan
gorals can be found at elevations from 1000 m to 4000 m.
- Habitat Regions
- temperate
- terrestrial
- Terrestrial Biomes
- chaparral
- forest
- scrub forest
- mountains
Physical Description
Himalayan gorals are medium sized herbivores and are the smallest of the goat-antelopes
(subfamily
Caprinae
). They range in body length from 81 to 130 cm and stand 56 to 80 cm tall. Himalayan
gorals weigh from 25 to 30 kg. They are stout animals; a strong and stocky build is
probably advantageous for maneuvering the craggy terrain of the Himalayas. Male and
females are similar in size and both genders have short, sharp horns that curve posteriorly.
Horns are rarely longer than 15 cm. Himalayan gorals are dark grey or brown with a
darker colored dorsal stripe and a lighter patch of hair on the throat. The hair is
short and coarse and males have manes from their necks to their chests.
Himalayan gorals lack a pre-orbital gland, which is present in closely related
serows
.
- Other Physical Features
- endothermic
- homoiothermic
- bilateral symmetry
- Sexual Dimorphism
- sexes alike
- ornamentation
Reproduction
Himalayan gorals are polygynous. Dominant males have mating rights to all females
in their ranges during the breeding season. This dominance is established through
threatening displays and combat with other males. When Himalayan gorals fight, they
attempt to wound the flank of their opponent with their small, dagger-like horns instead
of engaging in head-to-head butting.
Males court females with low stretches, lip curling, spraying of urine, and tail raising.
- Mating System
- polygynous
Himalayan gorals mate in November and December so that young are born in spring and
early summer when vegetation is abundant. Females give birth to one young per breeding
season. Gestation lasts roughly 6 months, after which the female gives birth in isolation.
After several days of hiding, the young begins to follow the mother. Juvenile gorals
are weaned 4 to 5 months after birth. They are sexually mature at 2 to 3 years of
age.
- Key Reproductive Features
- iteroparous
- seasonal breeding
- gonochoric/gonochoristic/dioecious (sexes separate)
- sexual
- fertilization
- viviparous
Female Himalayan gorals raise their young with no help from the males, as males only
associate with herds during the breeding season. Young Himalayan gorals follow their
mother until weaned and remain with the group to which their mother belongs after
reaching sexual maturity.
- Parental Investment
- precocial
- female parental care
-
pre-fertilization
- provisioning
-
protecting
- female
-
pre-hatching/birth
-
provisioning
- female
-
protecting
- female
-
provisioning
-
pre-weaning/fledging
-
provisioning
- female
-
protecting
- female
-
provisioning
-
pre-independence
-
provisioning
- female
-
protecting
- female
-
provisioning
- post-independence association with parents
Lifespan/Longevity
The expected lifespan of Himalayan gorals, once they have survived to maturity, is
15 years.
Behavior
Himalayan gorals are diurnal but most active during the early morning and evening (crepuscular). After grazing in the early morning, the animals spend the day relaxing in caves or on cliffs. Himalayan gorals are very agile and can easily traverse the arduous landscape of the Himalayas. They make small altitudinal migrations seasonally; in the colder months they move to lower altitudes to graze, returning to higher altitudes during warmer months. When snow blankets all available food sources, Himalayan gorals push the snow away with their snouts instead of digging or scratching with their hooves.
Himalayan gorals are gregarious, but adult males live in solitude until the breeding
season. Females and juveniles graze in herds of four to twelve individuals
- Key Behaviors
- cursorial
- terricolous
- diurnal
- crepuscular
- motile
- migratory
- sedentary
- solitary
- territorial
- social
- dominance hierarchies
Home Range
Herds of females and juveniles occupy a range of 40 hectares. Solitary males occupy
a 22 to 25 hectare territory.
Communication and Perception
Himalayan gorals rely on their vision and hearing to sense their surroundings. Their
acute sense of sight allows them to see predators while they rest or graze. These
animals use a series of snorts, whistles, and sneezes to indicate alarm to other gorals.
Like most other mammals, it is likely that olfaction is an important sense as well.
- Other Communication Modes
- pheromones
Food Habits
Himalayan gorals are herbivores and eat a wide range of vegetation including herbs,
shoots, roots, twigs, lichen, fungi, leaves, and grasses.
- Plant Foods
- leaves
- roots and tubers
- wood, bark, or stems
- seeds, grains, and nuts
- flowers
- bryophytes
- lichens
- Other Foods
- fungus
Predation
Snow leopards
,
Eurasian lynx
,
wolves
,
wild dogs
, and
humans
all prey on Himalayan gorals. The dark grey pelage of Himalayan gorals and their
relatively sedentary behavior during the day allow them to blend in with the surrounding
mountainside. Their agility in rough terrain also helps them to avoid less sure-footed
predators.
- Anti-predator Adaptations
- cryptic
Ecosystem Roles
Himalayan gorals are grazers, impacting vegetation communities in their mountain habitats.
They also provide valuable source of nutrition for several predators that inhabit
the unforgiving crags of the Himalayas.
Economic Importance for Humans: Positive
Humans hunt Himalayan gorals for meat, wool, and hides. Goral blood is also used medicinally
in some Asian cultures, although its efficacy is not proven.
- Positive Impacts
- food
- body parts are source of valuable material
Economic Importance for Humans: Negative
There are no known adverse affects of Naemorhedus goral on humans.
Conservation Status
Himalayan gorals are listed as near threatened as of 2008 by the IUCN. They are also
on Appendix I of CITES. Populations are declining due to habitat loss and hunting.
As humans begin to construct roads that spread deeper into the mountains of the Himalayas,
populations of Himalayan gorals increasingly lose habitats that suit their isolated
nature. Additionally, advances in weapon technology allow hunters to kill gorals from
farther distances.
One study found that Himalayan gorals graze primarily on several grasses that are
endemic to the Himalayas. If Himalayan gorals become more seriously endangered, human
cultivation of these grasses could provide surviving Himalayan gorals with a reliable
and preferred food source.
Other Comments
Additional Links
Contributors
Eric Cohen (author), University of Michigan-Ann Arbor, Phil Myers (editor, instructor), Museum of Zoology, University of Michigan-Ann Arbor, Tanya Dewey (editor), Animal Diversity Web.
- Palearctic
-
living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- oriental
-
found in the oriental region of the world. In other words, India and southeast Asia.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- temperate
-
that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).
- terrestrial
-
Living on the ground.
- chaparral
-
Found in coastal areas between 30 and 40 degrees latitude, in areas with a Mediterranean climate. Vegetation is dominated by stands of dense, spiny shrubs with tough (hard or waxy) evergreen leaves. May be maintained by periodic fire. In South America it includes the scrub ecotone between forest and paramo.
- forest
-
forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.
- scrub forest
-
scrub forests develop in areas that experience dry seasons.
- mountains
-
This terrestrial biome includes summits of high mountains, either without vegetation or covered by low, tundra-like vegetation.
- endothermic
-
animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.
- bilateral symmetry
-
having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.
- sexual ornamentation
-
one of the sexes (usually males) has special physical structures used in courting the other sex or fighting the same sex. For example: antlers, elongated tails, special spurs.
- polygynous
-
having more than one female as a mate at one time
- iteroparous
-
offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).
- seasonal breeding
-
breeding is confined to a particular season
- sexual
-
reproduction that includes combining the genetic contribution of two individuals, a male and a female
- fertilization
-
union of egg and spermatozoan
- viviparous
-
reproduction in which fertilization and development take place within the female body and the developing embryo derives nourishment from the female.
- young precocial
-
young are relatively well-developed when born
- female parental care
-
parental care is carried out by females
- diurnal
-
- active during the day, 2. lasting for one day.
- crepuscular
-
active at dawn and dusk
- motile
-
having the capacity to move from one place to another.
- migratory
-
makes seasonal movements between breeding and wintering grounds
- sedentary
-
remains in the same area
- solitary
-
lives alone
- territorial
-
defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement
- social
-
associates with others of its species; forms social groups.
- dominance hierarchies
-
ranking system or pecking order among members of a long-term social group, where dominance status affects access to resources or mates
- visual
-
uses sight to communicate
- acoustic
-
uses sound to communicate
- pheromones
-
chemicals released into air or water that are detected by and responded to by other animals of the same species
- visual
-
uses sight to communicate
- tactile
-
uses touch to communicate
- acoustic
-
uses sound to communicate
- chemical
-
uses smells or other chemicals to communicate
- cryptic
-
having markings, coloration, shapes, or other features that cause an animal to be camouflaged in its natural environment; being difficult to see or otherwise detect.
- food
-
A substance that provides both nutrients and energy to a living thing.
- herbivore
-
An animal that eats mainly plants or parts of plants.
- folivore
-
an animal that mainly eats leaves.
References
Duckworth, J., J. MacKinnon. 2008. "Naemorhedus Goral" (On-line). Red List of Endangered Species. Accessed April 05, 2009 at http://www.iucnredlist.org .
Fukhar-i-Abbas, F., T. Akhtar, A. Mian. 2008. Food & Feeding Preferences of Himalayan Gray Goral in Pakistan and Arad Jammu and Kashmir. Zoo Biology , 27/5: 371-380.
Geist, V. 1971. Mountain Sheep: A Study in Behavior & Evolution . Chicago: University of Chicago Press.
Matthews, L. 1971. Life of Mammals . New York: Universe Books.
Morris, D. 1965. The Mammals . New York: Harper & Row Publishing.
Thomas, W. 2001. Endangered Wildlife and Plants of the World . Marshall Cavendish. Accessed April 10, 2009 at http://books.google.com/books?id=40jA0MOWejIC&pg=PA640&lpg=PA640&dq=goral+predators&source=bl&ots=ZcMKdjReUS&sig=QaUtyiqpZ_u0aXiEOOt8jbiPnhU&hl=en&ei=2ozfSdKkBIrinQfh89mkCQ&sa=X&oi=book_result&ct=result&resnum=3#PPA640,M1 .
2003. Caprinae. Pp. 87-98 in Grzinek's Animal Life Encyclopedia , Vol. 16, 2 Edition. Detroit: Thomson-Gale Publishing.
1987. Naemorhedus Goral. The Encyclopedia of Mammals , Vol. 1, 1 Edition. New York, NY: Facts on File Publishing.