Geographic Range
        In North America, this species breeds as far north as Alaska and the Arctic coast
            of Canada south into the Great Lakes region and westward across Idaho, Nevada and
            Oregon.  It also breeds in the extreme southeastern United States and Cuba. The winter
            range of this species includes parts of California, New Mexico, Arizona, Texas, Georgia,
            Florida and northern Mexico.  Populations of sandhill cranes are also found in northeastern
            Siberia, Andyrland, and on the Chyukotski peninsula and Wrangel Island.
        
- Biogeographic Regions
- nearctic
- palearctic
- neotropical
Habitat
Most sandhill crane populations nest in open grasslands, such as wet meadows, and freshwater marshes or bogs. There are six subspecies of sandhill cranes, and each typically nests in the open, wet grassland habitats of their region. The Cuban population of sandhill cranes ( Grus canadensis nesiotes ) is an exception, inhabiting dry, isolated regions, sometimes in rocky and mountainous terrain.
        Sandhill cranes prefer to be far from human habitation.  However, during migration,
            they are commonly seen feeding on crops and crop residue in agricultural fields. 
            At night they congregate to roost in large marshes.
        
- Habitat Regions
- temperate
- tropical
- terrestrial
- Terrestrial Biomes
- savanna or grassland
- forest
- Other Habitat Features
- agricultural
- riparian
Physical Description
Sandhill cranes are large birds with heavy bodies and long necks and legs. They stand about 1.2 m tall, with wing spans of about 2 m. They are uniformly grayish, with a white cheek and a bald red crown. Sandhill cranes can be distinguished from other large wading birds in flight by their outstretched neck, and their wingbeats, which are a slow downward beat followed by a quick upward flick.
Male and female sandhill cranes are similar in appearance, though males are usually larger than females. For example, males of the G. c. canadensis subspecies average 3.75 kg whereas females average 3.34 kg. Juvenile sandhill cranes tend to be much more brownish than adults.
        There are six subspecies of sandhill cranes.  These subspecies are distinguished based
            upon morphology (size) and geographic separation.
        
- Other Physical Features
- endothermic
- bilateral symmetry
- polymorphic
- Sexual Dimorphism
- male larger
Reproduction
        Sandhill cranes are perennially monogamous.  Breeding pairs remain together from year
            to year, maintaining the pair bond by performing courtship displays, remaining in
            close proximity and calling together in unison.  Breeding pairs form during spring
            migration.  This species is noted for its elaborate courtship displays.  Five courtship
            displays have been identified as part of "dancing," the primary mechanism of pair
            formation in this species.  These displays are the Upright wing stretch, Horizontal
            head pump, Bow, Vertical leap and Vertical toss.  Three courtship displays are used
            exclusively by paired adults to maintain the pair bond and synchronize reproductive
            development.  These are the Bill up, Copulation and Unison call displays.
        
- Mating System
- monogamous
Sandhill cranes raise one brood per year. In non-migratory populations, egg-laying can begin as early as December or as late as August. In migratory populations, egg-laying usually begins between early April and late May. Both members of a breeding pair build the nest using plant material from the surrounding areas. Nest sites are usually in marshes, bogs, or swales, though cranes will occasionally nest on dry land.
The female lays 1 to 3 (usually 2) eggs that are oval-shaped and dull brown with reddish brown markings. Both parents participate in incubation, which lasts 29 to 32 (usually 30) days. Incubation begins with the laying of the first egg and continues until the second egg has hatched. The chicks are precocial; they hatch covered in down, with their eyes open and are able to leave the nest within 24 hours of hatching. The parents brood the chicks for up to 3 weeks after hatching. They feed the young intensively for the first few weeks, and with decreasing frequency until they reach independence at 9 or 10 months old.
        The chicks remain with their parents until 1 or 2 months before the parents begin
            laying the next clutch of eggs.  After leaving their parents, the chicks form nomadic
            flocks with other subadults and non-breeders.  They remain with these flocks until
            they form breeding pairs and begin breeding between the ages of 2 and 7 years old.
        
- Key Reproductive Features
- iteroparous
- seasonal breeding
- gonochoric/gonochoristic/dioecious (sexes separate)
- sexual
- oviparous
        Sandhill cranes provide extended biparental care to their young.  Both members of
            a breeding pair build the nest, incubate the eggs and feed and protect the chicks
            for up to 10 months after hatching.
        
- Parental Investment
- precocial
- 
         
          pre-fertilization
         
         - provisioning
- 
           
            protecting
           
           - female
 
 
- 
         
          pre-hatching/birth
         
         - 
           
            protecting
           
           - male
- female
 
 
- 
           
            protecting
           
           
- 
         
          pre-weaning/fledging
         
         - 
           
            provisioning
           
           - male
- female
 
- 
           
            protecting
           
           - male
- female
 
 
- 
           
            provisioning
           
           
- 
         
          pre-independence
         
         - 
           
            provisioning
           
           - male
- female
 
- 
           
            protecting
           
           - male
- female
 
 
- 
           
            provisioning
           
           
- post-independence association with parents
Lifespan/Longevity
Sandhill cranes that reach independence are expected to live about seven years. The oldest known sandhill crane lived at least 21.6 years.
Behavior
Sandhill cranes are diurnal and partially migratory. Northern populations move south during the winter months whereas southern populations remain near the breeding sites year round.
        Cranes are usually found in pairs and family groups.  During the migration and winter,
            family groups may join with non-mated cranes to form survival groups that feed and
            roost together.  These survival groups often congregate at migratory staging areas
            and on the wintering grounds.
        
Home Range
Two studies resident sandhill crane populations in Florida estimated average home ranges of 657 and 1366 hectares.
Communication and Perception
        Sandhill cranes communicate primarily using physical displays (see Mating Systems
            section) and vocalizations.  Adult sandhill cranes have a repertoire of more than
            a dozen calls, which can be described as variations on "trills", "purrs" and "rattles".
            Calls are used in territorial advertisement, social interactions and to notify others
            of a nearby predator.  Breeding pairs may call in duet in order to advertise their
            occupancy of a territory.
        
Food Habits
        Sandhill cranes are omnivorous birds that use their bills to probe for subsurface
            food and glean seeds and other foods.  These birds feed on land or in shallow marshes
            with vegetation.  Foods vary depending on what is available. Cultivated grains such
            as corn, wheat and sorghum are a major food source in their diet when available. 
            In northern latitudes, a wider variety of foods are consumed, including berries, small
            mammals, insects, snails, reptiles, and amphibians.
        
- Primary Diet
- omnivore
- Animal Foods
- mammals
- amphibians
- reptiles
- insects
- mollusks
- Plant Foods
- seeds, grains, and nuts
- fruit
Predation
        When approached by an avian predator, sandhill cranes fly at the predator, kicking
            it with their feet.  When approached by a mammalian predator, sandhill cranes move
            toward the predator with their wings spread and their bill pointed at the predator.
            If the predator persists, sandhill cranes will attack, hissing, stabbing with their
            bills and kicking with their feet.
        
Ecosystem Roles
        Young and sick sandhill cranes provide food for their predators.  Sandhill cranes
            affect the populations of species that they prey upon.  They also host at least 24
            different species of parasites.
        
Economic Importance for Humans: Positive
        Sandhill cranes feed on insects and rodents that may damage crops.
        
- Positive Impacts
- controls pest population
Economic Importance for Humans: Negative
Sandhill cranes feed on crops where they are available.
- Negative Impacts
- crop pest
Conservation Status
        Sandhill cranes are protected under the U.S. Migratory Bird Act and CITES Appendix
            II.  Two subspecies of sandhill cranes,
        
         Grus canadensis nesiotes
        
        (Cuba sandhill crane) and
        
         Grus canadensis pulla
        
        (Mississippi sandhill crane), are federally endangered in the United States.  Low
            reproduction rates limit population recovery in this species, especially by  the mid-continent
            population, which is subject to hunting.  Reintroduction of captive-reared birds has
            been instrumental in maintaining population size.   Protection of wetland habitats
            is also essential for the survival of this species.
        
Additional Links
Contributors
Kari Kirschbaum (author, editor), Animal Diversity Web.
Marie S. Harris (author), University of Michigan-Ann Arbor.
- Nearctic
- 
          living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.   
- native range
- 
          the area in which the animal is naturally found, the region in which it is endemic. 
- Palearctic
- 
          living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.   
- introduced
- 
          referring to animal species that have been transported to and established populations in regions outside of their natural range, usually through human action. 
- Neotropical
- 
          living in the southern part of the New World. In other words, Central and South America.   
- native range
- 
          the area in which the animal is naturally found, the region in which it is endemic. 
- temperate
- 
          that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle). 
- tropical
- 
          the region of the earth that surrounds the equator, from 23.5 degrees north to 23.5 degrees south. 
- terrestrial
- 
          Living on the ground. 
- tropical savanna and grassland
- 
          A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia. 
- savanna
- 
          A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome. 
- temperate grassland
- 
          A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands. 
- forest
- 
          forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality. 
- marsh
- 
          marshes are wetland areas often dominated by grasses and reeds. 
- swamp
- 
          a wetland area that may be permanently or intermittently covered in water, often dominated by woody vegetation. 
- bog
- 
          a wetland area rich in accumulated plant material and with acidic soils surrounding a body of open water. Bogs have a flora dominated by sedges, heaths, and sphagnum. 
- agricultural
- 
          living in landscapes dominated by human agriculture. 
- riparian
- 
          Referring to something living or located adjacent to a waterbody (usually, but not always, a river or stream). 
- endothermic
- 
          animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds. 
- bilateral symmetry
- 
          having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria. 
- polymorphic
- 
          "many forms." A species is polymorphic if its individuals can be divided into two or more easily recognized groups, based on structure, color, or other similar characteristics. The term only applies when the distinct groups can be found in the same area; graded or clinal variation throughout the range of a species (e.g. a north-to-south decrease in size) is not polymorphism. Polymorphic characteristics may be inherited because the differences have a genetic basis, or they may be the result of environmental influences. We do not consider sexual differences (i.e. sexual dimorphism), seasonal changes (e.g. change in fur color), or age-related changes to be polymorphic. Polymorphism in a local population can be an adaptation to prevent density-dependent predation, where predators preferentially prey on the most common morph. 
- monogamous
- 
          Having one mate at a time. 
- iteroparous
- 
          offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes). 
- seasonal breeding
- 
          breeding is confined to a particular season 
- sexual
- 
          reproduction that includes combining the genetic contribution of two individuals, a male and a female 
- oviparous
- 
          reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body. 
- young precocial
- 
          young are relatively well-developed when born 
- diurnal
- 
          - active during the day, 2. lasting for one day.
 
- motile
- 
          having the capacity to move from one place to another. 
- migratory
- 
          makes seasonal movements between breeding and wintering grounds 
- sedentary
- 
          remains in the same area 
- territorial
- 
          defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement 
- social
- 
          associates with others of its species; forms social groups. 
- visual
- 
          uses sight to communicate 
- acoustic
- 
          uses sound to communicate 
- omnivore
- 
          an animal that mainly eats all kinds of things, including plants and animals 
- visual
- 
          uses sight to communicate 
- tactile
- 
          uses touch to communicate 
- acoustic
- 
          uses sound to communicate 
- chemical
- 
          uses smells or other chemicals to communicate 
References
Peterson, R. 1980. Eastern Birds; A completely new field guide to all the birds of eastern and central North America . New York: Houghton Mifflin Company.
Tacha, T., S. Nesbitt, P. Vohs. 1992. Sandhill Crane ( Grus canadensis ). Pp. 1-24 in The Birds of North America , Vol. 31. Philadelphia, PA: The Academy of Natural Sciences; Washington, DC: The American Ornithologists' Union.
