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area above the spring high tide line) of sand beaches, from the water line up to the dunes. Ocypode quadrata is small, having a carapace length of about 5 cm (2 inches) at maturity. They are either straw-colored or grayish-white. They have a quadrate carapace, large club-shaped eyestalks, unequal chelipeds (claws) and long walking legs. Males are generally larger than females. After hatching from an egg, Ocypode quadrata has five zoea stages and one megalopa stage. The megalopa stage requires at least 35