Aquila rapaxtawny eagle

Ge­o­graphic Range

Tawny ea­gles occur from Ro­ma­nia east to south­ern Rus­sia and Mon­go­lia, and south through India and much of Africa. (Camp­bell, 1983)

Habi­tat

Tawny ea­gles favor arid cli­mates but oc­cupy a wide range of habi­tats in­clud­ing deserts, steppes, open sa­van­nah, open grass­land, moun­tain­ous re­gions, and cul­ti­vated steppes. The tend to avoid dense forests. (Brown, et al., 1982; Bur­ton, 1983; Camp­bell, 1983; Chan­ning, 2006)

  • Range elevation
    600 to 4500 m
    1968.50 to 14763.78 ft

Phys­i­cal De­scrip­tion

Aquila rapax plumage varies from very dark brown to light brown shades with black­ish flight feath­ers and tail, light col­ored stripes or bars on the wings, and a pale lower back. Tawny ea­gles with darker shades of brown gen­er­ally have tawny col­oration on the body, dis­tin­guish­ing them from sim­i­lar species of ea­gles, which lack any tawny col­oration. The eyes are brown and the beak is yel­low with a dark tip. Fe­males are typ­i­cally larger than males, oth­er­wise the sexes are sim­i­lar. Im­ma­ture tawny ea­gles are paler and more streaked than are adults, some­times "blonde" (white). Tawny ea­gles were thought to be syn­ony­mous with steppe ea­gles (Aquila ni­palen­sis). Steppe ea­gles are larger than tawny ea­gles (up to twice the size in weight) and are darker in color. (Brown, 1977; Brown, et al., 1982; Camp­bell, 1983; Clancey, 1964; Maclean, 1988)

  • Sexual Dimorphism
  • female larger
  • Range mass
    1950 to 2500 g
    68.72 to 88.11 oz
  • Range length
    65 to 72 cm
    25.59 to 28.35 in
  • Range wingspan
    1.72 to 1.85 m
    5.64 to 6.07 ft

Re­pro­duc­tion

Tawny ea­gles are monog­a­mous, pair­ing for life. Be­hav­ior prior to and dur­ing mat­ing varies for this species, but usu­ally in­volves un­du­lat­ing dis­plays made by the male fol­lowed by mu­tual soar­ing dis­plays. Epigamic dis­play, dis­play that oc­curs dur­ing breed­ing, may in­volve high cir­cling, alone or in pairs, over the nest­ing site. The male may per­form a se­ries of "pot hooks" which in­volves a se­ries of grad­ual dives and swoops, with lit­tle to no wing flap­ping. The fe­male may turn over and pre­sent her claws in re­sponse to the male swoop­ing over her. Males and fe­males may lock claws in flight. Ac­tual mat­ing usu­ally oc­curs at or near the nest­ing site. (Brown and Amadon, 1968; Brown, 1977; Brown, et al., 1982; Maclean, 1988)

Tawny ea­gles breed once yearly. Breed­ing sea­son varies by ge­o­graphic lo­ca­tion, but typ­i­cally oc­curs from April to July. Males and fe­males both build the nest. Males col­lect nest ma­te­r­ial, while fe­males as­sem­ble the nest. Nests are usu­ally in trees or tele­phone poles and are oc­cu­pied for one to three years be­fore they are aban­doned. Fe­male tawny ea­gles lay one to three eggs at three day in­ter­vals and in­cu­bate them for ap­prox­i­mately 45 days. Al­though males pri­mar­ily feed off­spring while they are young, both par­ents bring food dur­ing fledg­ing, which oc­curs at ap­prox­i­mately 76 to 85 days old. Tawny ea­gles begin to fly around ten weeks, but chicks re­main in the nest for ap­prox­i­mately 5 more weeks after their first flight and re­main re­liant on their par­ents for food dur­ing this pe­riod. After that, the young be­come in­de­pen­dent. Sib­lings are ag­gres­sive to­wards one an­other, many times re­sult­ing in the death of the younger hatch­ling, usu­ally within the first few days of hatch­ing. (Brown, et al., 1982; Bur­ton, 1983; Camp­bell, 1983; Knys­tau­tas, 1993; Maclean, 1988)

  • Breeding interval
    Tawny eagles breed once yearly.
  • Breeding season
    The breeding season of tawny eagles is generally from April to July.
  • Range eggs per season
    1 to 3
  • Range time to hatching
    39 to 45 days
  • Range fledging age
    76 to 85 days
  • Average fledging age
    84 days
  • Average time to independence
    120 days
  • Range age at sexual or reproductive maturity (female)
    3 to 4 years
  • Range age at sexual or reproductive maturity (male)
    3 to 4 years

Fe­male tawny ea­gles gen­er­ally in­cu­bate the eggs, oc­ca­sion­ally as­sisted by males. For the first ten days fe­males re­main by the nest, brood­ing day and night, and males may also brood or shade the chicks from the sun. After 7 days fe­males leave the chicks for ex­tended pe­ri­ods, but stay near the nest to pro­tect them. They con­tinue to perch near the nest for ap­prox­i­mately forty days. At fifty days, nei­ther males nor fe­males are near the nest dur­ing the day. Males brings most of the food for the chicks, but may be as­sisted by fe­males after fifty days. Chicks make their first flight around 84 days old and may re­main in the nest for up to forty days after the first flight. How­ever, some young re­main with the par­ents until the fol­low­ing breed­ing sea­son. (Brown, et al., 1982; Bur­ton, 1983; Camp­bell, 1983)

  • Parental Investment
  • altricial
  • pre-fertilization
    • provisioning
    • protecting
      • female
  • pre-hatching/birth
    • protecting
      • male
      • female
  • pre-weaning/fledging
    • provisioning
      • male
      • female
    • protecting
      • male
      • female
  • pre-independence
    • provisioning
      • male
      • female

Lifes­pan/Longevity

The lifes­pan of adult ea­gles is dif­fi­cult to de­ter­mine in the wild. The old­est golden ea­gles, also in the genus Aquila, are recorded to have lived for sixty years. The av­er­age lifes­pan of golden ea­gles is 18 years, and they live ap­prox­i­mately 40 to 45 years in cap­tiv­ity. The lifes­pan of tawny ea­gles may be sim­i­lar to these val­ues. In East Africa it is es­ti­mated that the lifes­pan of tawny ea­gles is 16 years on av­er­age, as­sum­ing that there is a 75% mor­tal­ity rate be­fore sex­ual ma­tu­rity. Ea­gles live much longer in cap­tiv­ity, rarely reach­ing the same ages in the wild due to the high mor­tal­ity rate within the first twelve months of life. (Brown, 1977; Brown, et al., 1982; Love, 1989; "Sedg­wick County Zoo", 2006)

  • Range lifespan
    Status: wild
    40.3 (high) years
  • Average lifespan
    Status: wild
    16 years
  • Typical lifespan
    Status: captivity
    40 to 45 years

Be­hav­ior

Tawny ea­gles are gre­gar­i­ous out­side of the breed­ing sea­son. Groups of at least twenty are com­monly ob­served, es­pe­cially when large amounts of food are lo­cally avail­able (such as swarms of lo­custs and large pop­u­la­tions of rats). Tawny ea­gles spend time near water where they can drink freely. Tawny ea­gles nor­mally fly only when ther­mals can as­sist in soar­ing, and usu­ally perch in trees dur­ing the day, es­pe­cially in heavy rain. Tawny ea­gles, like most other ea­gles, are di­ur­nal. Some pop­u­la­tions of tawny ea­gles are mi­gra­tory but many are not. (Brown, et al., 1982; Camp­bell, 1983; Chan­ning, 2006)

  • Range territory size
    25 to 100 km^2
  • Average territory size
    35-55 km^2

Home Range

Tawny ea­gles tend to oc­cupy the same ter­ri­tory for many years, some­times even decades, in pairs. Ter­ri­to­ries are usu­ally fairly far apart (20 km^2) to avoid food steal­ing by neigh­bor­ing tawny ea­gles. Ter­ri­to­ries may range from 25 to 100 km^2 in area, most are 35-55 km^2. (Brown, et al., 1982)

Com­mu­ni­ca­tion and Per­cep­tion

Tawny ea­gles are gen­er­ally fairly silent, ex­cept when ag­gra­vated or dis­play­ing. Their call can be de­scribed as a sharp kwok kwok. Oc­ca­sion­ally tawny ea­gles will call dur­ing acts of piracy. Fe­males may call from the nest, so­lic­it­ing food. In gen­eral, vi­sion is acute among ea­gles, and is likely to be their most im­por­tant sense. They are able to see prey clearly at dis­tances and up close. Their acute vi­sion may also help in es­tab­lish­ing ter­ri­to­ries. Hear­ing is also an im­por­tant sense for tawny ea­gles, as it helps them lo­cate prey when they are hunt­ing. (Brown, 1977; Brown, et al., 1982; Camp­bell, 1983)

Food Habits

Tawny ea­gles are gen­er­al­ist car­ni­vores, they will eat in­sects, car­rion, and small an­i­mals such as ro­dents. They are the only type of eagle that scav­enge from hu­mans reg­u­larly. Tawny ea­gles are also well known for rob­bing prey from other rap­tors, even birds much larger than they are. (Brown, 1977; Camp­bell, 1983)

  • Primary Diet
  • carnivore
    • eats terrestrial vertebrates
  • Animal Foods
  • mammals
  • carrion
  • insects

Pre­da­tion

Nest pre­da­tion by crows oc­curs more reg­u­larly in the nests of tawny ea­gles than in other species, pos­si­bly due to the open­ness of the nest site. There is very lit­tle other in­for­ma­tion on other preda­tors or be­hav­iors to pre­vent pre­da­tion in tawny ea­gles. Tawny ea­gles are large birds of prey once they reach adult­hood, they are prob­a­bly do not have many preda­tors. (Brown, et al., 1982)

  • Known Predators

Ecosys­tem Roles

Tawny ea­gles are pri­mary preda­tors. They are also pi­rat­i­cal and op­por­tunis­tic. They will steal prey from other an­i­mals as well as scav­enge al­ready dead prey. (Brown, 1977; Camp­bell, 1983)

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Tawny ea­gles are im­por­tant mem­bers of the ecosys­tems in which they live.

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

Tawny ea­gles are pi­rat­i­cal and steal and scav­enge food from other an­i­mals and hu­mans. They may oc­ca­sion­ally take do­mes­tic live­stock, al­though not often enough to have a sig­nif­i­cant neg­a­tive im­pact on hu­mans. (Brown and Amadon, 1968)

Con­ser­va­tion Sta­tus

The cur­rent IUCN Red List sta­tus of this species is "lower risk/least con­cern."

Con­trib­u­tors

Tanya Dewey (ed­i­tor), An­i­mal Di­ver­sity Web.

Joslyn West­phal (au­thor), Kala­ma­zoo Col­lege, Ann Fraser (ed­i­tor, in­struc­tor), Kala­ma­zoo Col­lege.

Glossary

Ethiopian

living in sub-Saharan Africa (south of 30 degrees north) and Madagascar.

World Map

Palearctic

living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.

World Map

acoustic

uses sound to communicate

agricultural

living in landscapes dominated by human agriculture.

altricial

young are born in a relatively underdeveloped state; they are unable to feed or care for themselves or locomote independently for a period of time after birth/hatching. In birds, naked and helpless after hatching.

arboreal

Referring to an animal that lives in trees; tree-climbing.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

carnivore

an animal that mainly eats meat

carrion

flesh of dead animals.

chemical

uses smells or other chemicals to communicate

desert or dunes

in deserts low (less than 30 cm per year) and unpredictable rainfall results in landscapes dominated by plants and animals adapted to aridity. Vegetation is typically sparse, though spectacular blooms may occur following rain. Deserts can be cold or warm and daily temperates typically fluctuate. In dune areas vegetation is also sparse and conditions are dry. This is because sand does not hold water well so little is available to plants. In dunes near seas and oceans this is compounded by the influence of salt in the air and soil. Salt limits the ability of plants to take up water through their roots.

diurnal
  1. active during the day, 2. lasting for one day.
endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

migratory

makes seasonal movements between breeding and wintering grounds

monogamous

Having one mate at a time.

motile

having the capacity to move from one place to another.

mountains

This terrestrial biome includes summits of high mountains, either without vegetation or covered by low, tundra-like vegetation.

native range

the area in which the animal is naturally found, the region in which it is endemic.

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

seasonal breeding

breeding is confined to a particular season

sedentary

remains in the same area

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

social

associates with others of its species; forms social groups.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

terrestrial

Living on the ground.

territorial

defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement

tropical savanna and grassland

A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.

savanna

A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.

temperate grassland

A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.

visual

uses sight to communicate

Ref­er­ences

Sedgewick County Zoo. 2006. "Sedg­wick County Zoo" (On-line). Golden Eagle. Ac­cessed No­vem­ber 09, 2006 at http://​www.​scz.​org/​animals/​e/​golden.​html.

Brown, L., E. Urban, K. New­man. 1982. The Birds of Africa Vol. 1. Lon­don: Aca­d­e­mic Press Inc.

Brown, L. 1977. Ea­gles of the World. New York, NY: Uni­verse Books.

Brown, L., D. Amadon. 1968. Ea­gles, Hawks, and Fal­cons of the World. New York, NY: Mc­Graw-Hill Book Com­pany.

Bur­ton, P. 1983. Van­ish­ing Ea­gles. New York, NY: Dodd, Mead & Com­pany, Inc.

Camp­bell, B. 1983. The Dic­tio­nary of Birds in Color. New York, NY: Ex­eter Books.

Chan­ning, K. 2006. "The Hawk Con­ser­vancy Trust" (On-line). Ac­cessed Oc­to­ber 15, 2006 at http://​www.​hawk-conservancy.​org/​priors/​frodo.​shtml.

Clancey, P. 1964. Birds of Natal and Zu­l­u­land. Lon­don: Oliver and Boyd Ltd.

Knys­tau­tas, A. 1993. Birds of Rus­sia. Lon­don: Harper Collins Pub­lish­ers.

Love, J. 1989. Ea­gles. Lon­don: Whit­tet Books Ltd.

Maclean, G. 1988. Robert's Birds of South­ern Africa. Lon­don: New Hol­land Pub­lish­ers Ltd.