Apus apuscommon swift

Ge­o­graphic Range

Com­mon swifts, Apus apus, can be found in al­most any re­gion from west­ern Eu­rope to east­ern Asia and from north­ern Scan­danavia and north­ern Siberia to North Africa, Hi­malayas, and cen­tral China. Apus apus can be found through­out this range dur­ing the breed­ing sea­son and, fol­low­ing mi­gra­tion, spends the win­ter months in South­ern Africa, from Zaire and Tan­za­nia south to Zim­babwe and Mozam­bique. ("Com­mon swift (Apus apus)", 1985; "Swifts", 2003; Ban­ner­man, 1955)

Habi­tat

The ma­jor­ity of the breed­ing habi­tat of A. apus is lo­cated in tem­per­ate zones, where there are suit­able trees for nest­ing and suf­fi­cient open spaces in which to fly to gather food. The habi­tat of Apus apus dur­ing the months fol­low­ing mi­gra­tion into Africa, how­ever, is trop­i­cal. Com­mon swifts have been ob­served breed­ing from sea level to sev­eral thou­sand me­ters in el­e­va­tion. Apus apus prefers areas with trees, or build­ings with open spaces, and is able to use ver­ti­cal sur­faces such as rock walls and chim­neys for nest­ing due to a unique phys­i­cal adap­ta­tion pos­sessed by all swifts (Apo­di­dae). ("Com­mon swift (Apus apus)", 1985; "Swifts", 2003; Bruun, et al., 1992; Ter­res, 1980)

Phys­i­cal De­scrip­tion

Com­mon swifts are 16 to 17 cm in length with a wingspan of 42 to 48 cm, de­pend­ing on the age of the in­di­vid­ual. Com­mon swifts are black-brown with the ex­cep­tion of a white to cream col­ored chin and throat (lo­cated di­rectly un­der­neath the beak). In ad­di­tion, the top­side of the flight feath­ers is paler brown-black in com­par­i­son to the rest of the body. Apus apus can also be dis­tin­guished by its mod­er­ately forked tail feath­ers, its nar­row, sickle-shaped wings, as well as its shrill, scream­ing call. Apus apus is fre­quently mis­taken for swal­lows. Apus apus is larger and has very a dif­fer­ent wing shape and flight pat­tern than do swal­lows. All species in the fam­ily Apo­di­dae pos­sess a unique mor­pho­log­i­cal char­ac­ter­is­tic, a lat­eral “grasp­ing foot” in which toes one and two are op­posed by toes three and four. This al­lows com­mon swifts to oc­cupy areas such as walls of rock, chim­neys, and other ver­ti­cal sur­faces that would be dif­fi­cult for other types of birds to in­habit. Apus apus is a sex­u­ally monomor­phic species, mean­ing that the males and fe­males look alike. There has been no sea­sonal or ge­o­graph­i­cal vari­a­tion re­ported in the ap­pear­ance. How­ever, it is pos­si­ble to dis­tin­guish ju­ve­niles from adults in the slight dif­fer­ence in rich­ness and uni­for­mity of their col­oration, as it is com­mon for ju­ve­niles to be blacker in color, as well as to have a pale fore­head, white-fringed feath­ers, and a starker white patch under the beak. This dis­tinc­tion is best ob­served at close range. ("Com­mon swift (Apus apus)", 1985; "Swifts", 2003; Bruun, et al., 1992; John­son, 1992)

  • Sexual Dimorphism
  • sexes alike
  • Average mass
    44.9 g
    1.58 oz
    AnAge
  • Average length
    16-17 cm
    in
  • Range wingspan
    42 to 48 cm
    16.54 to 18.90 in
  • Average basal metabolic rate
    0.4372 W
    AnAge

Re­pro­duc­tion

Apus apus usu­ally first breeds at two years of age, but the age of the first breed­ing can vary based upon the avail­abil­ity of nest­ing sites. The com­mon swift is a monog­a­mous species, they are faith­ful to nest­ing sites each year and to mates dur­ing a breed­ing sea­son. The male A. apus typ­i­cally chooses the nest site. Upon the ar­rival of the fe­male shortly there­after (usu­ally within a pe­riod of days), the nest­ing site is pro­tected by the pair. The nest is typ­i­cally com­posed of grass, leaves, hay, straw, and flower petals (among other things). The nest­ing site usu­ally in­cludes the nest it­self and the areas di­rectly sur­round­ing the nest. Courtship, some cop­u­la­tion, and the rear­ing of the chicks all occur at this site. Colonies of A. apus typ­i­cally in­clude 30 to 40 nest sites, re­flect­ing the gre­gar­i­ous na­ture of the com­mon swift mat­ing sys­tem. Apus apus is more likely to fight to de­fend a nest­ing site than it is to de­fend a mate. Males at­tract their fe­male part­ners through at­tain­ment of a good nest­ing site prior to their meet­ing. Upon their first meet­ing it is not un­usual for the ini­tial re­sponses of the po­ten­tial mates, both male and fe­male, to be hos­tile. If in­ter­ested and un­paired, the fe­male will enter the nest site ten­ta­tively, thereby invit­ing her po­ten­tial part­ner to stroke her chin with its bill. If this en­counter is suc­cess­ful, the fe­male may also in­vite her po­ten­tial part­ner to al­lo­preen. Al­lo­preen­ing is the process by which birds smooth or clean each oth­ers feath­ers with their beak or bill. This mu­tual ac­tion be­gins the pair-bond­ing process. ("Com­mon Swift (Apus apus)", 2001; "Swifts", 2003; Ban­ner­man, 1955)

Com­mon swifts typ­i­cally breed from late April to early May through mid-Sep­tem­ber when the young are fledged. One of the most unique char­ac­ter­is­tics of A. apus is its abil­ity to mate while in flight, al­though they also can mate while on the nest. Mat­ing oc­curs every few days fol­low­ing the ar­rival of suit­able weather, until a few days after the young have fledged. Fol­low­ing a suc­cess­ful cop­u­la­tion, any­where from one to four white eggs may be laid, how­ever a clutch size of two is most com­mon. Eggs must then be du­ti­fully in­cu­bated for 19 to 20 days while the em­bryos de­velop. Both par­ents par­tic­i­pate in the in­cu­ba­tion of the clutch. After the young hatch, it can take an ad­di­tional 27 to 45 days be­fore fledg­ing oc­curs, usu­ally more than 27 days. ("Com­mon Swift (Apus apus)", 2001; "Swifts", 2003; Ban­ner­man, 1955; John­son, 1992)

  • Breeding interval
    Apus apus breeds once yearly.
  • Breeding season
    Common swifts breed from late April to early May through mid-September
  • Average eggs per season
    2
  • Range time to hatching
    19 to 20 days
  • Range fledging age
    27 to 45 days
  • Average age at sexual or reproductive maturity (female)
    2 years
  • Average age at sexual or reproductive maturity (male)
    2 years

Both par­ents take turns in­cu­bat­ing the clutch fol­low­ing fer­til­iza­tion and prior to hatch­ing. For the du­ra­tion of the first week fol­low­ing hatch­ing, the clutch is typ­i­cally brooded all day long. Dur­ing the sec­ond week, the young are brooded for ap­prox­i­mately half of the day. For the re­main­der of the time, until the clutch is fledged, they are rarely brooded dur­ing the day, but are al­most al­ways cov­ered at night. Both par­ents par­tic­i­pate equally in all as­pects of the rais­ing of the young. In the event that un­usu­ally bad weather per­sists or food sources be­come scarce dur­ing the time shortly after the hatch­ing of the young, the young pos­sess the abil­ity to be­come semi-tor­pid, a hi­ber­na­tion-like state, thereby re­duc­ing the en­ergy de­mands of their rapidly grow­ing bod­ies. This adap­ta­tion al­lows young A. apus to sur­vive with lit­tle food for 10 to 15 days. Dur­ing the time from hatch­ing until fledg­ing, the young are fed al­most ex­clu­sively in the nest. The young are fed food-balls con­sist­ing of in­sects gath­ered by the par­ents dur­ing flight and held to­gether with a sali­vary gland prod­uct, cre­at­ing the food bolus. While the young are smaller, they will share a food bolus among them. How­ever, once young are larger they be­come able to swal­low an en­tire food bolus on their own. ("Com­mon Swift (Apus apus)", 2001; "Swifts", 2003; John­son, 1992)

  • Parental Investment
  • altricial
  • pre-fertilization
    • provisioning
    • protecting
      • female
  • pre-hatching/birth
    • provisioning
      • female
    • protecting
      • male
      • female
  • pre-weaning/fledging
    • provisioning
      • male
      • female
    • protecting
      • male
      • female

Lifes­pan/Longevity

Apus apus is typ­i­cally long-lived. A com­mon swift banded in Swe­den was re-trapped at the age of 17. The an­nual sur­vival rate for the adults is 65-83%. (; "Com­mon Swift (Apus apus)", 2001; Ter­res, 1980)

Be­hav­ior

Apus apus is a highly gre­gar­i­ous species. Com­mon swifts typ­i­cally nest, roost, mi­grate, and hunt for food in groups through­out the year. In ad­di­tion, they are unique in their abil­ity to stay aloft for ex­tended pe­ri­ods of time. It is not un­com­mon for these swifts to spend the en­tire day on the wing, only land­ing to feed young or to roost at night. It is es­ti­mated that com­mon swifts fly at least 560 miles per day dur­ing the nest­ing sea­son, il­lus­trat­ing their en­durance and strength, as well tremen­dous aer­ial abil­ity. They can also mate and for­age for food while in the air. Com­mon swifts typ­i­cally fly in lower air­space when the weather is poor (cold, windy, and/or wet), and will move to higher air­spaces when the weather is more fa­vor­able for ex­tended aer­ial ac­tiv­ity. (; "Swifts", 2003; Bruun, et al., 1992; John­son, 1992)

Home Range

Apus apus has been known to con­sis­tently for­age more than 48-61 km from nest­ing or roost­ing sites.

Com­mu­ni­ca­tion and Per­cep­tion

Com­mu­ni­ca­tion be­tween Apus apus oc­curs al­most ex­clu­sively through the use of dif­fer­ent vo­cal­iza­tions, or calls, and by changes in body lan­guage. The types of calls used by A. apus are largely de­pen­dent upon its age. There are dif­fer­ent calls used by the adults than by the young. The most com­mon call dur­ing flight, a long, shrill ‘sreee’, is used in in­nu­mer­able con­texts by adults. Also among the vo­cal­iza­tions of the adults are those given dur­ing al­lo­preen­ing (nest-call), those fol­low­ing de­feat in a fight (pip­ing-call), as well as those pre­ced­ing cop­u­la­tion (pre-cop­u­la­tory call). The most com­mon call used by young is the food-call, used to beg for food from a par­ent upon its re­turn. (; "Swifts", 2003; Bruun, et al., 1992; Svens­son and Grant, 1999)

Food Habits

Com­mon swifts are in­sec­tiv­o­rous, feed­ing solely on aer­ial in­sects and spi­ders that it gath­ers in its mouth as it glides through the air. The in­sects are gath­ered to­gether in­side the throat through the use of a prod­uct from the sali­vary glands, to form a food-ball or bolus. Apus apus is com­monly at­tracted to swarms of in­sects, as it aides in the ease of col­lect­ing suf­fi­cient food. It has been es­ti­mated that there are an av­er­age of 300 in­sects per bolus. These num­bers may vary based upon the abun­dance of prey. Among some of the most com­monly con­sumed in­sects are aphids (Hemiptera), wasps, bees, and ants (Hy­menoptera), bee­tles (Coleoptera), and flies (Diptera). ("Com­mon Swift (Apus apus)", 2001; "Swifts", 2003; John­son, 1992; Svens­son and Grant, 1999)

  • Animal Foods
  • insects
  • terrestrial non-insect arthropods

Pre­da­tion

Most no­table among the anti-preda­tor adap­ta­tions of A. apus is its aer­ial mas­tery, al­low­ing these birds to avoid most of their nat­ural preda­tors, in­clud­ing Eurasian hob­bies (Falco sub­bu­teo), spar­rowhawks (Ac­cip­iter nisus), and buz­zards (Buteo buteo), by tak­ing to the air. In ad­di­tion, the choice of nest­ing sites on ver­ti­cal sur­faces such as rock walls and chim­neys makes it dif­fi­cult for com­mon swifts to be preyed upon be­cause of the level of dif­fi­culty as­so­ci­ated with ac­cess­ing the nest area. The plain col­oration of Apus apus also is ad­van­ta­geous for preda­tor eva­sion as it makes them dif­fi­cult to see when they are not in the air. (; "Com­mon Swift (Apus apus)", 2001; "Swifts", 2003)

Ecosys­tem Roles

As a preda­tor, A. apus con­tributes to the con­trol of the in­sect pop­u­la­tion. (; Ban­ner­man, 1955; John­son, 1992)

Com­men­sal/Par­a­sitic Species
  • mites (Acari)
  • lice (Anoplura)

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Be­sides the fact that the com­mon swift fre­quently nests in close as­so­ci­a­tion with hu­mans, A. apus has no sig­nif­i­cant eco­nom­i­cal im­pact on hu­mans. Apus apus may offer a slight ben­e­fit to hu­mans by con­sum­ing pest in­sects such as mos­qui­tos. How­ever, it is un­likely that A. apus it­self would have a sig­nif­i­cant im­pact on these pest pop­u­la­tions. In some places com­mon swifts are en­cour­aged to nest in man­made struc­tures so that the young can be har­vested for food, how­ever, this prac­tice is not very com­mon. Also the nests of some swifts are used by in­dige­nous peo­ples of Asia as the key in­gre­di­ent in bird’s nest soup. (; "Com­mon Swift (Apus apus)", 2001; "Swifts", 2003)

  • Positive Impacts
  • food
  • controls pest population

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

Com­mon swifts have no no­table neg­a­tive im­pacts upon hu­mans, with the ex­cep­tion of the oc­ca­sional nui­sance of hav­ing them nest in the eaves and open spaces in the rooftops of many cities and vil­lages across Eu­rope. (; "Swifts", 2003; Bruun, et al., 1992; John­son, 1992; Svens­son and Grant, 1999)

  • Negative Impacts
  • household pest

Con­ser­va­tion Sta­tus

Apus apus is nei­ther threat­ened nor en­dan­gered. (; "Swifts", 2003)

Con­trib­u­tors

Tanya Dewey (ed­i­tor), An­i­mal Di­ver­sity Web.

Katie Thomp­son (au­thor), Kala­ma­zoo Col­lege, Ann Fraser (ed­i­tor, in­struc­tor), Kala­ma­zoo Col­lege.

Glossary

Ethiopian

living in sub-Saharan Africa (south of 30 degrees north) and Madagascar.

World Map

Palearctic

living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.

World Map

acoustic

uses sound to communicate

altricial

young are born in a relatively underdeveloped state; they are unable to feed or care for themselves or locomote independently for a period of time after birth/hatching. In birds, naked and helpless after hatching.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

carnivore

an animal that mainly eats meat

chemical

uses smells or other chemicals to communicate

diurnal
  1. active during the day, 2. lasting for one day.
endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

food

A substance that provides both nutrients and energy to a living thing.

forest

forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.

insectivore

An animal that eats mainly insects or spiders.

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

migratory

makes seasonal movements between breeding and wintering grounds

monogamous

Having one mate at a time.

motile

having the capacity to move from one place to another.

native range

the area in which the animal is naturally found, the region in which it is endemic.

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

seasonal breeding

breeding is confined to a particular season

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

social

associates with others of its species; forms social groups.

suburban

living in residential areas on the outskirts of large cities or towns.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

terrestrial

Living on the ground.

tropical

the region of the earth that surrounds the equator, from 23.5 degrees north to 23.5 degrees south.

urban

living in cities and large towns, landscapes dominated by human structures and activity.

visual

uses sight to communicate

Ref­er­ences

2001. Com­mon Swift (Apus apus). C El­ph­ick, J Dun­ning, D Sib­ley, eds. Na­tional Audubon So­ci­ety Sib­ley Guide to Bird Life and Be­hav­ior. New York: Al­fred a Knopf, Inc..

1985. Com­mon swift (Apus apus). Pp. 657-670 in D Snow, ed. The Birds of the West­ern Palearc­tic: Birds of Eu­rope, the Mid­dle East, and North Africa. New York: Ox­ford Uni­ver­sity Press..

2003. Swifts. Pp. 421-425, 429-430 in M Hutchins, ed. Grz­imek’s An­i­mal Life En­cy­clo­pe­dia, Sec­ond Edi­tion, Vol. 9. New York: The Gale Group, Inc..

Ban­ner­man, D. 1955. Order Apod­i­formes, Sub-Or­der Apodes, Fam­ily Apo­di­dae, Genus Apus. Pp. 1-12 in The Birds of the British Isles, Vol. IV, 1 Edi­tion. Ed­in­burgh: Oliver and Boyd Ltd..

Bruun, B., H. Delin, L. Svens­son. 1992. Birds of Britain and Eu­rope. Lon­don: Ham­lyn Pub­lish­ing Group Lim­ited.

John­son, L. 1992. Birds of Eu­rope with North Africa and the Mid­dle East. Lon­don: A & C Black.

Svens­son, L., P. Grant. 1999. Birds of Eu­rope. Prince­ton: Prince­ton Uni­ver­sity Press.

Ter­res, J. 1980. Swift Fam­ily. Pp. 868-870 in The Audubon So­ci­ety En­cy­clo­pe­dia of North Amer­i­can Birds. New York: Al­fred A Knopf, Inc..