Ensis directus

Ge­o­graphic Range

Ensis di­rec­tus is found along the At­lantic coast from Canada to South Car­olina. It lives in the in­ter­tidal zone or sub­ti­dal zone in the sand or muddy bot­toms.

(Jobin and Jobin 1997, Gos­ner 1978)

Habi­tat

Ensis di­rec­tus lives in the sandy bot­toms in the in­ter­tidal or sub­ti­dal zones along the At­lantic coast. It is usu­ally found in colonies. It is not mi­gra­tory and there­fore it re­mains in its habi­tat year round.

(Gos­ner 1978)

Phys­i­cal De­scrip­tion

Ensis di­rec­tus has a thin, elon­gated shell that is slightly curved. It ranges from yel­low­ish to dark brown in color. The length of E. di­rec­tus is about six times its width. It can grow to be about 10 in. It has a coat­ing around its shell to pro­tect it from erod­ing in the mud or sand. Ensis di­rec­tus is a bi­valve, which means that its shell has two parts. The body of Ensis di­rec­tus is sur­rounded by the man­tle and the man­tle is seper­ated into two parts. Each part of the man­tle se­cretes a shell. The two shells are con­nected by an elas­tic lig­a­ment that al­lows for it to open and close. Both parts are usu­ally iden­ti­cal and are made up of cal­cium car­bon­ate and pro­tein. It has a huge foot that al­lows it to move through water or to bur­row in the sand. When the foot is ex­tended all the way, it is al­most as long as the clam's body.

(Alexan­der 1979; Lipp­son 1984)

Re­pro­duc­tion

There are sep­a­rate male and fe­male sexes in Ensis di­rec­tus. The males re­lease their sperm into the water and the sperm en­ters the fe­male through open­ings. The eggs are fer­til­ized in the in­te­rior of the gill by the sperm and these newly fer­til­ized zy­gotes de­velop into larva. This larva is then re­leased into the sur­round­ing water. There are two lar­val stages. The first stage is the tro­cophore stage that has small lar­vae that are free swim­ming. They are pear shaped, translu­cent, and cil­i­ated. The sec­ond stage is the veliger stage, which is also a free-swim­ming lar­val stage. It has a very long pelagic or plank­ton stage, which means that the lar­vae float freely within the water. This al­lows for the lar­vae to spread over large dis­tances. This larva then set­tles onto the sand or mud and be­gins to de­velop into an adult. The body will de­velop as well as the man­tle. The man­tle will then se­crete and line the shell.

(Kinder­s­ley 2001; Ogden 2001)

Be­hav­ior

Ensis di­rec­tus bur­rows into the sand using its foot and it only sur­faces dur­ing high tide. It will climb close to the sur­face so that only its siphons are ex­posed. These two siphons are used for fil­ter­ing food and water. When low tide oc­curs, it bur­rows back down below the sur­face. E. di­rec­tus is an ex­tremly fast bur­rower and is very hard to be caught due to its great speed. It is also a re­mark­able swim­mer. It is able to pro­pel it­self through water by ex­pelling water through its shell and draw­ing in its foot. This ac­tion is re­peated over and over again al­low­ing it to move through the water.

(Alexan­der 1979; Cooper 1960)

Food Habits

Ensis di­rec­tus is a fil­ter feeder that fil­ters water through its shell in order to ob­tain food. When feed­ing, E. di­rec­tus stays very close to the sur­face and its siphons are stick­ing up through the sur­face. The water is drawn into the shell through the man­tle cav­ity by cilia. These cilia cover the cteni­dia, or gills, in the clam. It passes along the gills and com­bines with mu­cous. The food is now trapped and the cilia drive the food into the di­ges­tive tract.

(Jobin and Jobin 1997)

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Ensis di­rec­tus is a very fast bur­rower and very dif­fi­cult to catch while it is still alive. How­ever, when it is caught it can be sold and eaten like many other types of clams. E. di­rec­tus is in sea­son dur­ing the months of July-Sep­tem­ber.

(Great North­ern Prod­ucts. 2001; Cooper 1960)

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

Un­known. The Jack­knife clam bur­rows deep and sur­faces only to ob­tain food and water. There does not seem to be any neg­a­tive ef­fect of the clam on hu­mans.

Con­ser­va­tion Sta­tus

Con­trib­u­tors

Kim­berly Cam­ponelli (au­thor), West­ern Mary­land Col­lege, Louise a. Paquin (ed­i­tor), West­ern Mary­land Col­lege.

Glossary

Atlantic Ocean

the body of water between Africa, Europe, the southern ocean (above 60 degrees south latitude), and the western hemisphere. It is the second largest ocean in the world after the Pacific Ocean.

World Map

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

coastal

the nearshore aquatic habitats near a coast, or shoreline.

ectothermic

animals which must use heat acquired from the environment and behavioral adaptations to regulate body temperature

native range

the area in which the animal is naturally found, the region in which it is endemic.

Ref­er­ences

Alexan­der, R. 1979. The In­ver­te­brates. Great Britain: Cam­bridge Uni­ver­sity Press.

Cooper, E. 1960. Sci­ence on the Shores and Banks. New York: Har­court, Brace and Com­pany.

Gos­ner, K. 1978. At­lantic Seashore. New York: Houghton Mif­flin Com­pany.

Great North­ern Prod­ucts, 2001. "Mol­luscs and Crus­ta­ceous" (On-line). Ac­cessed April 27, 2001 at http://​northernproducts.​com/​html/​molluscs___​crustaceous.​html.

Jobin, A., R. Jobin. 1997. "The As­sateague Nat­u­ral­ist" (On-line). Ac­cessed April 20, 2001 at http://​www.​assateague.​com.

Kinder­s­ley, D. "Mol­lusca - Anatomy" (On-line). Ac­cessed May 9, 2001 at http://​www.​chardikalaa.​com/​~gurtej/hmwk/bio/mol­lusca/.

Lipp­son, J. 1984. Life in the Chesa­peake Bay. Bal­ti­more: Johns Hop­kins Uni­ver­sity Press.

Ogden, M. "Mol­lusca" (On-line). Ac­cessed May 9, 2001 at http://​www.​personal.​psu.​edu/​users/​m/​a/​mao153/​.