Jacana spinosanorthern jacana

Ge­o­graphic Range

North­ern ja­canas (Ja­cana spin­osa) range from Mex­ico to Panama, al­though they are some­times seen into the south­ern United States. They occur mainly in coastal areas. In Mex­ico they are found from Tamauli­pas on Gulf of Cal­i­for­nia coast and Sinaloa on the Pa­cific coast south to west­ern Panama. North­ern ja­canas are also found in the An­til­lean is­lands of Cuba, His­pan­iola, Ja­maica, and the West In­dies. (Jenni and Mace, 1999; Kauf­man, 1996)

Habi­tat

North­ern ja­canas in­habit many types of wet­lands in­clud­ing; ponds, marshes and lake mar­gins. Pond edges with abun­dant float­ing veg­e­ta­tion are fa­vored. North­ern ja­canas oc­ca­sion­ally for­age in wet grassy areas and flooded fields. (Jenni and Mace, 1999; Kauf­man, 1996)

  • Aquatic Biomes
  • lakes and ponds
  • rivers and streams
  • temporary pools
  • coastal

Phys­i­cal De­scrip­tion

North­ern ja­canas are medium sized wad­ing birds with long legs and ex­tremely elon­gated toes. Adults are rel­a­tively dark over­all; they have a black neck, head and breast. The back, un­der­sides, and tail are dark ru­fous. They have a yel­low bill with a white base and a yel­low shield on the fore­head. Ju­ve­niles are bi-col­ored with a white un­der­side and a darker back, head, and neck. Ju­ve­niles have a white su­per­cil­ium and white lores. Both adults and ju­ve­niles have con­spic­u­ous yel­low flight feath­ers that are vis­i­ble when they raise their wings. North­ern ja­canas av­er­age 241 mm in length with a wingspan av­er­ag­ing 508 mm. Males were found to be sig­nif­i­cantly smaller than fe­males. A study from Costa Rica showed that fe­males weighted 145.4g and males weighed 86.9 on av­er­age. (Jenni and Col­lier, 1972; Sib­ley, 2003)

  • Sexual Dimorphism
  • female larger
  • Range mass
    86.9 to 145.4 g
    3.06 to 5.12 oz
  • Average length
    241 mm
    9.49 in
  • Average wingspan
    508 mm
    20.00 in

Re­pro­duc­tion

The breed­ing sys­tem in north­ern ja­canas is un­usual and is an ex­am­ple of polyandry. Both males and fe­males will de­fend ter­ri­to­ries against other mem­bers of the same sex. Fe­males mate with up to four in­di­vid­ual males and lay eggs as fre­quently as every nine days. Males may be polyg­a­mous when new fe­males ar­rive. When north­ern ja­canas breed, males build plat­forms that are used for so­lic­i­ta­tion dis­plays and cop­u­la­tion. The plat­forms are then used as nests. The fe­male or the male may so­licit to each other and this be­hav­ior leads to cop­u­la­tion. So­lic­i­ta­tion is done by call­ing or pos­tur­ing and it can be ini­ti­ated by the male or the fe­male. When the fe­male as­sumes a pre-cop­u­la­tory po­si­tion, the male may fly up 10 me­ters or more be­fore fly­ing back down, land­ing on her back or land­ing along­side and hop­ping on her back. The male may ini­ti­ate a mat­ing event by as­sum­ing the pre-cop­u­la­tory po­si­tion and the fe­male then joins him. Most cop­u­la­tion at­tempts are un­suc­cess­ful. When they are suc­cess­ful, the male gives dis­tinct calls. So­lic­i­ta­tion and cop­u­la­tion occur less often once a male has eggs in his nest, but may occur again when the young are 4 to 5 weeks old. (Jenni and Col­lier, 1972; Sib­ley, 2001)

Al­though north­ern ja­canas are ca­pa­ble of breed­ing all year long, they usu­ally breed at the be­gin­ning of the rainy sea­son. Dur­ing that time, fe­males mate with up to four males and lay eggs as fre­quently as every nine days. Males may be polyg­a­mous when new fe­males ar­rive, but extra pair cop­u­la­tions are not known to occur when the fe­male holds her ter­ri­tory. When north­ern ja­canas breed, males build plat­forms that are used for so­lic­i­ta­tion dis­plays and cop­u­la­tion. The plat­forms are then used as nests. As in many other mem­bers of the order Charadri­iformes, four eggs are typ­i­cally laid per clutch. The male typ­i­cally sits on the eggs and adds nest­ing ma­te­r­ial in the form of aquatic plants pe­ri­od­i­cally. The fe­male may visit the nest site on oc­ca­sion and shade the eggs. The eggs hatch in around 28 days. The young leave the nest around 24 hours after hatch­ing and will fol­low the male to good for­ag­ing spots. Be­cause fe­males may have sev­eral mates, some or all of the young that he raises may not be his. The de­vel­op­ment time in north­ern ja­canas is slow; young fledge in around 8 weeks and may stay in their natal ter­ri­tory for more than 12 months. (Jenni and Col­lier, 1972; Jenni and Mace, 1999)

  • Breeding interval
    Time between breeding efforts in males is not known.
  • Breeding season
    Breeding corresponds with the start of the rainy season, which varies throughout the species range.
  • Average eggs per season
    4
  • Average time to hatching
    28 days
  • Average fledging age
    57.2 days
  • Range time to independence
    12 (low) months

In north­ern ja­canas, males ex­clu­sively con­struct nests, in­cu­bate eggs, and care for the young. Fe­males do not help to raise off­spring gen­er­ally, but may pro­vide some de­fense against preda­tors. Males build a nest on the plat­form that was used for so­lic­i­ta­tion and cop­u­la­tion. After the fe­male lays eggs in the nest the male con­tin­ues to build the nest by fling­ing ma­te­r­ial over his shoul­der in the di­rec­tion of the nest site. Brood­ing is mainly done in the morn­ing and evening. Dur­ing rainy con­di­tions, cool tem­per­a­tures and times of in­creased cloud cover, the amount of time spent brood­ing in­creases. The young leave the nest at about 24 hours after hatch­ing. At that point the male leads the nestlings to good for­ag­ing sites and con­tin­ues to pro­tect them. (Betts and Jenni, 1991; Jenni and Mace, 1999; Sib­ley, 2001)

  • Parental Investment
  • pre-fertilization
    • provisioning
    • protecting
      • female
  • pre-hatching/birth
    • provisioning
      • female
    • protecting
      • male
  • pre-weaning/fledging
    • provisioning
      • male
    • protecting
      • male
      • female
  • pre-independence
    • provisioning
      • male
    • protecting
      • male
      • female

Lifes­pan/Longevity

Lit­tle is known about lifes­pan in north­ern ja­canas. One male held his ter­ri­tory for 6.5 years and had to be around 2 years old to have a ter­ri­tory in the first place. (Jenni and Mace, 1999)

Be­hav­ior

North­ern ja­canas are highly so­cial and fre­quently occur in small colonies with a dom­i­nant fe­male and 1 to 4 males. North­ern ja­canas are di­ur­nal and spend much of their time walk­ing on emer­gent veg­e­ta­tion and search­ing for aquatic in­sects and seeds. They pick food off of the sur­face of plants, the sur­face of the water, or just below the water. Their feet are highly spe­cial­ized for this for­ag­ing be­hav­ior. Their small size and large feet allow these birds to ex­ploit this for­ag­ing habi­tat, as they can eas­ily sup­port their weight on sur­face aquatic veg­e­ta­tion. (Jenni and Col­lier, 1972; Kauf­man, 1996)

Home Range

North­ern ja­canas have spe­cific home ter­ri­to­ries that they use for breed­ing and feed­ing. Male ter­ri­to­ries av­er­aged 0.36 acres and fe­male ter­ri­to­ries av­er­aged 0.88 acres at one site in Costa Rica. Fe­male ter­ri­to­ries con­tain be­tween one and four males, which they breed with. Fe­males help in main­tain­ing the ter­ri­to­ries of the males by in­ter­ven­ing on be­half of the res­i­dent male when there is a ter­ri­to­r­ial dis­pute with an­other male. If the fe­male loses her ter­ri­tory to an­other fe­male, then the new­comer will have ac­cess to the males in that ter­ri­tory. Male ter­ri­to­ries may last con­sid­er­ably longer than fe­male ter­ri­to­ries. A male in Costa Rica held a ter­ri­tory for 6.5 years and the own­er­ship by fe­males changes 10 times in the same time pe­riod. (Jenni and Col­lier, 1972; Jenni and Mace, 1999)

Com­mu­ni­ca­tion and Per­cep­tion

North­ern ja­canas com­mu­ni­cate with vi­sual dis­plays and calls. Re­peated-note calls are given in a wide va­ri­ety of sit­u­a­tions. Notes with high am­pli­tude, note length, and du­ra­tion are used when the bird is under phys­i­cal at­tack or when it can't fend off an in­trud­ing con­spe­cific. The na­ture of the re­peated note calls say a lot about the in­ten­tion of the caller. North­ern ja­canas also com­mu­ni­cate by giv­ing groups of notes. This is most often given by males when his chicks are near. Flight calls are also given. Threat dis­plays are given by out­stretch­ing the wings and point­ing them for­ward. Males may do a sub­mis­sive dis­play in the pres­ence of fe­males in which they crouch down. Dur­ing the sub­mis­sive dis­play the male swings his head lat­er­ally while the fe­male pecks at the base of his neck. (Jenni and Mace, 1999)

Food Habits

North­ern ja­canas eat what­ever in­sects they can glean off of aquatic plant sur­faces. They turn over float­ing plants using their feet and bills while search­ing for in­sects hid­ing on the un­der-sur­face (Sib­ley 2001).They also eat flow­ers that are opened by pur­ple gallinules (Por­phyrio mar­tinica) . Adults will oc­ca­sion­ally take small fish. In a closely re­lated species, wat­tled ja­canas (Ja­cana ja­cana), the diet was made up of 20% seeds, with most seeds from spikerush (Eleocharis species), and 80% in­ver­te­brates, mainly in­sects. (Jenni and Col­lier, 1972; Os­borne and Bourne, 1977)

  • Animal Foods
  • fish
  • insects
  • aquatic crustaceans
  • Plant Foods
  • seeds, grains, and nuts
  • flowers

Pre­da­tion

Pos­si­ble preda­tors in­clude boa con­stric­tors (Boa con­stric­tor), spec­ta­cled caimans (Caiman croc­o­dilus), snap­ping tur­tles (Chely­dra ser­pentina) and var­i­ous rap­tors and mam­mals. Pur­ple gallinules (Por­phyrio mar­tinica) are the most com­mon preda­tors of north­ern ja­cana eggs and off­spring, they will take them out of nests when the male is away. For this rea­son, north­ern ja­canas will at­tack pur­ple gallinules when they see them in their ter­ri­tory. At­tacks con­sist of aer­ial at­tacks and charg­ing with wings spread on the sub­strate. At­tacks are ac­com­pa­nied by re­peated-note calls. (Jenni and Mace, 1999; Stephens, 1984)

Ecosys­tem Roles

North­ern ja­canas are pre­sum­ably im­por­tant sec­ondary con­sumers in the marshy habi­tats they in­habit. At a 2.8 hectare pond in Costa Rica be­tween 20 and 25 ja­canas were pre­sent at a given time. Given the fact that they are rel­a­tively small, en­dother­mic an­i­mals, they pre­sum­ably have a high caloric re­quire­ment. Be­cause of this, ja­canas prob­a­bly con­trol in­sect pop­u­la­tions. North­ern ja­canas also con­tribute to the en­er­getic needs of pur­ple gallinules (Por­phyrio mar­tinica). (Jenni and Mace, 1999)

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Al­though north­ern ja­canas pro­vide no di­rect ben­e­fits to hu­mans, they are cer­tainly a source of en­joy­ment for nat­u­ral­ist and bird­ers.

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

There are no known ad­verse ef­fects of north­ern ja­canas on hu­mans.

Con­ser­va­tion Sta­tus

North­ern ja­canas ap­pear to be com­mon through­out most of their range, but could be­come vul­ner­a­ble with loss of wet­lands. (Kauf­man, 1996)

Con­trib­u­tors

Tanya Dewey (ed­i­tor), An­i­mal Di­ver­sity Web.

Ryne Ruther­ford (au­thor), North­ern Michi­gan Uni­ver­sity, Alec R. Lind­say (ed­i­tor, in­struc­tor), North­ern Michi­gan Uni­ver­sity.

Glossary

Nearctic

living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.

World Map

Neotropical

living in the southern part of the New World. In other words, Central and South America.

World Map

acoustic

uses sound to communicate

agricultural

living in landscapes dominated by human agriculture.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

carnivore

an animal that mainly eats meat

chemical

uses smells or other chemicals to communicate

coastal

the nearshore aquatic habitats near a coast, or shoreline.

diurnal
  1. active during the day, 2. lasting for one day.
dominance hierarchies

ranking system or pecking order among members of a long-term social group, where dominance status affects access to resources or mates

ecotourism

humans benefit economically by promoting tourism that focuses on the appreciation of natural areas or animals. Ecotourism implies that there are existing programs that profit from the appreciation of natural areas or animals.

endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

freshwater

mainly lives in water that is not salty.

insectivore

An animal that eats mainly insects or spiders.

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

marsh

marshes are wetland areas often dominated by grasses and reeds.

motile

having the capacity to move from one place to another.

native range

the area in which the animal is naturally found, the region in which it is endemic.

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

polyandrous

Referring to a mating system in which a female mates with several males during one breeding season (compare polygynous).

riparian

Referring to something living or located adjacent to a waterbody (usually, but not always, a river or stream).

seasonal breeding

breeding is confined to a particular season

sedentary

remains in the same area

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

social

associates with others of its species; forms social groups.

suburban

living in residential areas on the outskirts of large cities or towns.

swamp

a wetland area that may be permanently or intermittently covered in water, often dominated by woody vegetation.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

terrestrial

Living on the ground.

territorial

defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement

tropical

the region of the earth that surrounds the equator, from 23.5 degrees north to 23.5 degrees south.

visual

uses sight to communicate

Ref­er­ences

Betts, B., D. Jenni. 1991. Time bud­gets and the adap­tive­ness of polyandry in north­ern ja­canas. Wil­son Bul­letin, 103: 578-597.

Jenni, D., G. Col­lier. 1972. Polyandry in the Amer­i­can Ja­cana. The Auk, 89: 743-765.

Jenni, D., T. Mace. 1999. "The North­ern Ja­cana (Ja­cana spin­osa), The birds of North Amer­ica On­line" (On-line). Ac­cessed April 08, 2008 at http://​bna.​birds.​cornell.​edu/​bna/​species/​467.

Kauf­man, K. 1996. Lives of North Amer­i­can birds. Boston, NewYork: Houghton Mif­flin Com­pany.

Os­borne, D., G. Bourne. 1977. Breed­ing Be­hav­ior and Food habits or the Wat­tled Ja­cana. The Con­dor, 79: 98-105.

Sib­ley, D. 2003. The Sib­ley Guide To Birds of East­ern North Amer­ica. New York: Al­fred A. Knopf.

Sib­ley, D. 2001. The Sib­ley Guide to Bird Life and Be­hav­ior. New York: Al­fred. A Knopf.

Stephens, M. 1984. In­ter­spe­cific Ag­gres­sive be­hav­ior of the Polyan­drous North­ern Ja­cana (Ja­cana spin­osa). The Auk, 101: 508-518.