Lagenorhynchus obliquidensPacific white-sided dolphin

Ge­o­graphic Range

Pa­cific white-sided dol­phins, Lagenorhynchus obliqi­u­dens, can be found through­out the tem­per­ate wa­ters of the north­ern Pa­cific Ocean. This pelagic species is en­demic to the cold re­gions of the Pa­cific, most com­monly oc­cur­ring be­tween the lat­i­tudes of 38 de­grees N and 47 de­grees N. The species in not found in arc­tic and trop­i­cal wa­ters.

They also range in the west­ern Pa­cific Ocean from the South Bering Sea to south­ern Japan.

White-sided dol­phins are found along the west coast of North Amer­ica, and mi­grate sea­son­ally in north-south (lat­i­tu­di­nal) di­rec­tions. These dol­phins are most abun­dant in shelf wa­ters off south­ern Cal­i­for­nia dur­ing the win­ter and off Ore­gon and Wash­ing­ton dur­ing late spring. His­tor­i­cally, Pa­cific white-sided dol­phins have been ob­served as far south as the Gulf of Cal­i­for­nia. In re­cent years, Lagenorhynchus obliqi­u­dens has de­clined in the Gulf of Cal­i­for­nia as water tem­per­a­tures have in­creased. (Sal­vadeo, et al., 2010; Stew­art, et al., 2002)

Habi­tat

Pa­cific white-sided dol­phins are com­mon in tem­per­ate seas. They are most abun­dant in deeper wa­ters in the open ocean and can dive as deep as 1000 m. Pa­cific white-sided dol­phins ex­ploit a large habi­tat in­clud­ing open ocean and near-shore wa­ters. ("Food of the Pa­cific white-sided dol­phin, Lagenorhynchus obliq­uidens, Dall's por­poise, Pho­coenoides dalli, and north­ern fur seal, Cal­lorhi­nus ursi­nus, off Cal­i­for­nia and Wash­ing­ton, with ap­pen­dices of size and food of Dall's por­poise from Alaskan wa­ters", 1980; Black, 1994; Brown and Nor­ris, 1956; Rech­steiner, et al., 2013; Ridg­way and Har­ri­son, 1998; Sal­vadeo, et al., 2010)

  • Range depth
    0 to 1000 m
    0.00 to 3280.84 ft

Phys­i­cal De­scrip­tion

Pa­cific white-sided dol­phins have a com­plex color pat­tern. They are black or dark gray with dis­tinct light gray stripes on their sides, dor­sal fin, and flip­pers. The stripes on their sides begin on the side of the face ahead of the eye and ex­tend to the base of the tail. A large patch of light gray dom­i­nates the an­te­rior of the dor­sal fin. Their eyes and mouth have dark col­oration. Dis­tin­guish­ing fea­tures of Pa­cific white-sided dol­phins are their tall, hooked, bi­col­ored dor­sal fins, lo­cated at the cen­ter of their back. Pa­cific white-sided dol­phins have pec­toral fins that share the same col­oration and are rounded.

White-sided dol­phins are 1.7-2.5 m long, with an av­er­age of 2.0 m. Males can reach this length of 2.5 m while fe­males can reach just 2.3 m. Adult white-sided dol­phins typ­i­cally weigh about 135-180 kg but males can weigh up to 200 kg. New­born Pa­cific white-sided dol­phins are 0.9-1.05 m long and weigh ap­prox­i­mately 15 kg.

The bod­ies of Pa­cific white-side dol­phins are ro­bust, and tor­pedo-shaped. Pa­cific white-sided dol­phins have 21 to 33 pairs of slightly-pointed teeth per jaw. Pa­cific white-sided dol­phins have short beaks. Their skulls have an av­er­age length of 39 cm. ("Food of the Pa­cific white-sided dol­phin, Lagenorhynchus obliq­uidens, Dall's por­poise, Pho­coenoides dalli, and north­ern fur seal, Cal­lorhi­nus ursi­nus, off Cal­i­for­nia and Wash­ing­ton, with ap­pen­dices of size and food of Dall's por­poise from Alaskan wa­ters", 1980; Brown and Nor­ris, 1956; Marino, 1999; Nowak, 2003; Pre­vost, 1996; Rech­steiner, et al., 2013; Ridg­way and Har­ri­son, 1998; Stew­art, et al., 2002)

  • Sexual Dimorphism
  • male larger
  • Range mass
    78 to 200 kg
    171.81 to 440.53 lb
  • Range length
    1.7 to 2.5 m
    5.58 to 8.20 ft
  • Average length
    2.0 m
    6.56 ft

Re­pro­duc­tion

Pa­cific white-sided dol­phins breed sea­son­ally from Au­gust to Oc­to­ber. Al­though very lit­tle is known about their mat­ing sys­tem, Con­nor (1994) sug­gests it is un­likely they are monog­a­mous. (Black, 1994; Con­nor, 1994; Fer­rero and Walker, 1996; Stew­art, et al., 2002)

The ges­ta­tion pe­riod of Pa­cific white-sided dol­phins lasts for 9-12 months. Most calv­ing oc­curs dur­ing late spring and sum­mer, and just one off­spring is born. Pa­cific white-sided dol­phins are 0.9 m-1.5 m in length at birth. Calves weigh, on av­er­age, 15 kg (range 13-22 kg) when born, and nurse for up to 18 months after birth. Pa­cific white-sided dol­phins give birth at a min­i­mum of once every two years.

Fe­male Pa­cific white-sided dol­phins reach sex­ual ma­tu­rity around a length of 1.7 m and be­tween the ages of 8 and 10 years. Males reach sex­ual ma­tu­rity at lengths around 1.7-1.8 m be­tween the ages of 10 and 11 years. Like other mam­mals, fer­til­iza­tion oc­curs in­ter­nally. Pa­cific white-sided dol­phins give birth to rel­a­tively well-de­vel­oped young. (Black, 1994; Fer­rero and Walker, 1996; Mann, 2000; Nowak, 2003; Rech­steiner, et al., 2013; Sal­vadeo, et al., 2010; Stew­art, et al., 2002)

  • Breeding interval
    Pacific white-sided dolphins breed once yearly
  • Breeding season
    August-October
  • Range number of offspring
    1 (high)
  • Average number of offspring
    1
  • Average number of offspring
    1
    AnAge
  • Range gestation period
    9 to 12 months
  • Range weaning age
    18 (high) months
  • Average weaning age
    12 months
  • Range time to independence
    18 (high) months
  • Range age at sexual or reproductive maturity (female)
    8 to 11 years
  • Range age at sexual or reproductive maturity (male)
    10 to 11 years

A Pa­cific white-sided dol­phin calf stays with its mother until it reach in­de­pen­dence around 18 months. Males are not in­volved in parental care be­yond the mat­ing process. (Fer­rero and Walker, 1996; Mann, 2000)

  • Parental Investment
  • precocial
  • female parental care
  • pre-weaning/fledging
    • provisioning
      • female
    • protecting
      • female
  • pre-independence
    • provisioning
      • female
    • protecting
      • female

Lifes­pan/Longevity

Weigl (2005) es­ti­mates Pa­cific white-sided dol­phins can live up to 46 years in cap­tiv­ity. Wil­son and Ruff (1999) re­ported a cap­tive Pa­cific white-sided dol­phin lived to be 36 years. It is un­known how long they can live in the wild. (Weigl, 2005; Wil­son and Ruff, 1999)

  • Range lifespan
    Status: captivity
    46 (high) years
  • Typical lifespan
    Status: captivity
    36 (high) years

Be­hav­ior

Pa­cific white-sided dol­phins are mo­bile crea­tures spe­cial­ized for swim­ming. They can be found in groups rang­ing from just a few in­di­vid­u­als to 100 in­di­vid­u­als. These groups are some­times sep­a­rated into smaller sub­groups or pods, which av­er­age about 15 mem­bers (typ­i­cally 10 to 25). Pods con­sist of all ages and both sexes.

Pa­cific white-sided dol­phins are so­cial an­i­mals. They are often found in mixed-species ag­gre­ga­tions with other cetaceans, pin­nipeds, and sea birds. They as­so­ci­ate most often with north­ern right whale dol­phins (Lis­sodel­phis bo­re­alis) and Risso’s dol­phins (Gram­pus griseus).

Pa­cific white-sided dol­phins are ac­tive at the water’s sur­face. They are known to ride the bow waves of ships. Pa­cific white-sided dol­phins are also well known for their aer­ial dis­plays. They leap clear of the water, and are the only dol­phins of the east­ern Pa­cific known to turn com­plete som­er­saults under nat­ural con­di­tions. (Black, 1994; Dahlheim, et al., 2009; Nowak, 2003; Ridg­way and Har­ri­son, 1998; Stew­art, et al., 2002)

Home Range

Gen­er­ally, most pelagic mam­mals will not de­fend or use a home range. No home range has been re­ported and it's as­sumed Pa­cific white-sided dol­phins do not have one.

Com­mu­ni­ca­tion and Per­cep­tion

Lagenorhynchus obliq­uidens uses echolo­ca­tion clicks that range pri­mar­ily from 20 to over 100 kHz. A study per­formed on cap­tive Pa­cific white-sided dol­phins re­vealed they use echolo­ca­tion clicks and touch to in­ter­pret their sur­round­ings. Pa­cific white-sided dol­phins also pro­duce whis­tle vo­cal­iza­tions, sim­i­lar to other dol­phins, in order to com­mu­ni­cate with one an­other. They com­mu­ni­cate vi­su­ally, acousti­cally, and through touch. Ac­cord­ing to Hen­der­son (2010) Pa­cific white-sided dol­phin vo­cal­iza­tions dif­fer across be­hav­ioral states. ("Food of the Pa­cific white-sided dol­phin, Lagenorhynchus obliq­uidens, Dall's por­poise, Pho­coenoides dalli, and north­ern fur seal, Cal­lorhi­nus ursi­nus, off Cal­i­for­nia and Wash­ing­ton, with ap­pen­dices of size and food of Dall's por­poise from Alaskan wa­ters", 1980; Baird, 1998; Black, 1994; Hen­der­son, 2010; Ridg­way and Har­ri­son, 1998; Yeater, et al., 2014)

Food Habits

Pa­cific white-sided dol­phins are ver­sa­tile and op­por­tunis­tic feed­ers. Their car­niv­o­rous diet con­sists of small fish found in large schools and squid (Loligo). They typ­i­cally con­sume Pa­cific her­ring (Clu­pea pal­lasii), capelin (Mal­lo­tus vil­lo­sus), mar­ket squid (Loligo opalescens), Pa­cific sar­dines (Sardinops sagax) and mack­erel (Scomber japon­i­cus) when avail­able. An av­er­age-sized Pa­cific white-sided dol­phin needs to con­sume around 12.5-15.8 kg of fish per day or ap­prox­i­mately 16-20% of its body mass daily.

Lagenorhynchus obliq­uidens has been ob­served feed­ing in groups av­er­ag­ing in size from 11-50 in­di­vid­u­als. They for­age while si­mul­ta­ne­ously milling. They are not con­sid­ered deep divers and typ­i­cally feed close to the shelf in shal­low depths. ("Food of the Pa­cific white-sided dol­phin, Lagenorhynchus obliq­uidens, Dall's por­poise, Pho­coenoides dalli, and north­ern fur seal, Cal­lorhi­nus ursi­nus, off Cal­i­for­nia and Wash­ing­ton, with ap­pen­dices of size and food of Dall's por­poise from Alaskan wa­ters", 1980; Black, 1994; Brown and Nor­ris, 1956; Dai­ley, et al., 1993; Rech­steiner, et al., 2013; Ridg­way and Har­ri­son, 1998; Stew­art, et al., 2002)

  • Animal Foods
  • fish
  • mollusks

Pre­da­tion

Due to their harm­ful ef­fects on fish­eries, hu­mans (Homo sapi­ens) kill Pa­cific white-sided dol­phins. In re­gions of Japan, they are hunted for their meat. Killer whales (Orch­i­nus orca) are also known preda­tors of Lagenorhynchus obliq­uidens. Both hu­mans and killer whales are preda­tors of adult and young Pa­cific white-sided dol­phins.

The col­oration of Pa­cific white-sided dol­phins al­lows them to be cam­ou­flaged. By hav­ing a light col­ored un­der­side and a dark col­ored back, Pa­cific white-sided dol­phins can blend in with their sur­round­ings when being viewed by a preda­tor from both above and below. (Black, 1994; Nowak, 2003; Stew­art, et al., 2002)

  • Anti-predator Adaptations
  • cryptic
  • Known Predators

Ecosys­tem Roles

Pa­cific white-sided dol­phins have a vital ecosys­tem role as con­sumers of fish. Seabirds often fol­low Pa­cific white-sided dol­phins and con­sume fish the dol­phins do not eat.

They are hosts for in­ter­nal par­a­sites in­clud­ing: ne­ma­todes (Anisakis sim­plex and Cras­si­cauda), trema­todes (Na­sitrema glo­bi­cepha­lae), and ces­todes (Phyl­loboth­rium del­phini, Mono­rygma grimaldii, and Stro­bilo­cephalus tri­an­gu­larus). They are also hosts for ex­ter­nal par­a­sites called cir­ri­pedes (Xenobal­anus). (Dai­ley and Walker, 1978; Mar­tin, et al., 1970; Nowak, 2003; Stew­art, et al., 2002)

Com­men­sal/Par­a­sitic Species

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Pa­cific white-sided dol­phins are hunted for their meat by Japan­ese fish­er­men. They are im­por­tant in the en­ter­tain­ment and tourism in­dus­try. For ex­am­ple, these dol­phins are trained in ma­rine mam­mal parks to per­form tricks and are used to ed­u­cate at­ten­dees about dol­phins. It is also com­mon for Pa­cific white-sided dol­phins to be spot­ted dur­ing whale-watch­ing ex­pe­di­tions. (Nowak, 2003; Stew­art, et al., 2002)

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

Pa­cific white-sided dol­phins neg­a­tively af­fect com­mer­cial fish­eries. They com­pete with the fish­eries for fish and often be­come tan­gled in drift­nets. (Nowak, 2003; Stew­art, et al., 2002)

Con­ser­va­tion Sta­tus

The great­est threat to Pa­cific white-sided dol­phins is high-seas drift­nets used by com­mer­cial fish­eries. The United Na­tions’ pro­hi­bi­tion of high-seas drift­net fish­ing is the most sig­nif­i­cant at­tempt to con­serve Pa­cific white-sided dol­phins in­ter­na­tion­ally. The United States re­quires fish­eries in the Pa­cific to have acoustic warn­ing sig­nals to help pre­vent dol­phins from being caught in drift­nets. Al­though these sig­nals have de­creased the num­ber of dol­phins killed by drift­nets, an av­er­age of 5.9 Pa­cific white-sided dol­phins are caught and killed an­nu­ally. Still, this is an im­prove­ment from his­tor­i­cal deaths; Hobbs and Jones (1993) re­ported ap­prox­i­mately 100,000 Pa­cific white-sided dol­phins were killed be­tween 1970 and 1990 by high-seas drift­net.

Pa­cific white-sided dol­phins are har­pooned in areas of Japan for human con­sump­tion. Al­though spe­cific num­bers are un­known, it is likely im­pact on Pa­cific white-sided dol­phins due to Japan­ese har­poon­ing is nom­i­nal.

Pa­cific white-sided dol­phins are an Ap­pen­dix II species, ac­cord­ing to CITES. Per this Ap­pen­dix, per­mits are re­quired to trade Pa­cific white-sided dol­phins in­ter­na­tion­ally. Per­mits are only granted if the trade will not be detri­men­tal to the sur­vival of the species. Pa­cific white-sided dol­phins are not threat­ened, but if cur­rent trade poli­cies re­main they could be­come en­dan­gered. The IUCN Red List lists Pa­cific white-sided dol­phins as a species of “least con­cern.” Lagenorhynchus obliq­uidens has no spe­cial sta­tus ac­cord­ing to the US Fed­eral List and the State of Michi­gan list. (Ham­mond, et al., 2012; Hobbs and Jones, 1993)

Con­trib­u­tors

Tay­lor Lay­ton (au­thor), Rad­ford Uni­ver­sity, Karen Pow­ers (ed­i­tor), Rad­ford Uni­ver­sity, April Tin­gle (ed­i­tor), Rad­ford Uni­ver­sity, Emily Clark (ed­i­tor), Rad­ford Uni­ver­sity, Cari Mc­gre­gor (ed­i­tor), Rad­ford Uni­ver­sity, Jacob Vaught (ed­i­tor), Rad­ford Uni­ver­sity, Tanya Dewey (ed­i­tor), Uni­ver­sity of Michi­gan-Ann Arbor.

Glossary

Pacific Ocean

body of water between the southern ocean (above 60 degrees south latitude), Australia, Asia, and the western hemisphere. This is the world's largest ocean, covering about 28% of the world's surface.

World Map

acoustic

uses sound to communicate

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

carnivore

an animal that mainly eats meat

chemical

uses smells or other chemicals to communicate

coastal

the nearshore aquatic habitats near a coast, or shoreline.

colonial

used loosely to describe any group of organisms living together or in close proximity to each other - for example nesting shorebirds that live in large colonies. More specifically refers to a group of organisms in which members act as specialized subunits (a continuous, modular society) - as in clonal organisms.

cryptic

having markings, coloration, shapes, or other features that cause an animal to be camouflaged in its natural environment; being difficult to see or otherwise detect.

echolocation

The process by which an animal locates itself with respect to other animals and objects by emitting sound waves and sensing the pattern of the reflected sound waves.

ecotourism

humans benefit economically by promoting tourism that focuses on the appreciation of natural areas or animals. Ecotourism implies that there are existing programs that profit from the appreciation of natural areas or animals.

endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

female parental care

parental care is carried out by females

food

A substance that provides both nutrients and energy to a living thing.

molluscivore

eats mollusks, members of Phylum Mollusca

motile

having the capacity to move from one place to another.

natatorial

specialized for swimming

native range

the area in which the animal is naturally found, the region in which it is endemic.

nomadic

generally wanders from place to place, usually within a well-defined range.

pelagic

An aquatic biome consisting of the open ocean, far from land, does not include sea bottom (benthic zone).

piscivore

an animal that mainly eats fish

saltwater or marine

mainly lives in oceans, seas, or other bodies of salt water.

seasonal breeding

breeding is confined to a particular season

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

social

associates with others of its species; forms social groups.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

visual

uses sight to communicate

viviparous

reproduction in which fertilization and development take place within the female body and the developing embryo derives nourishment from the female.

young precocial

young are relatively well-developed when born

Ref­er­ences

Na­tional Oceanic and At­mos­pheric Ad­min­is­tra­tion. Food of the Pa­cific white-sided dol­phin, Lagenorhynchus obliq­uidens, Dall's por­poise, Pho­coenoides dalli, and north­ern fur seal, Cal­lorhi­nus ursi­nus, off Cal­i­for­nia and Wash­ing­ton, with ap­pen­dices of size and food of Dall's por­poise from Alaskan wa­ters. 86137. Seat­tle, Wash­ing­ton: NOAA, NMFS. 1980.

Baird, R. 1998. An in­ter­ac­tion be­tween Pa­cific white-sided dol­phins and a neona­tal har­bor por­poise. Mam­malia, 62/1: 129-133.

Black, N. 1994. Be­hav­ior and Ecol­ogy of Pa­cific White-Sided Dol­phins (Lagenorhynchus obliq­uidens) in Mon­terey Bay, Cal­i­for­nia (The­sis). San Fran­cisco, Cal­i­for­nia: San Fran­cisco State Uni­ver­sity.

Brown, D., K. Nor­ris. 1956. Ob­ser­va­tions of cap­tive and wild cetaceans. Jour­nal of Mam­mal­ogy, 37/3: 311-326.

Con­nor, R. 1994. The Lives of Whales and Dol­phins. New York: Henry Holt and Com­pany.

Dahlheim, M., P. White, J. Waite, G. Eck­ert. 2009. Cetaceans of south­east Alaska: Dis­tri­b­u­tional and sea­sonal oc­cur­rence. Jour­nal of Bio­geog­ra­phy, 36/3: 410-426.

Dai­ley, M., D. Reish, J. An­der­son. 1993. Ecol­ogy of the South­ern Cal­i­for­nia Blight: A Syn­the­sis and In­ter­pre­ta­tion. Oak­land, CA: Uni­ver­sity of Cal­i­for­nia Press.

Dai­ley, M., W. Walker. 1978. Par­a­sitism as a fac­tor (?) in sin­gle strand­ings of south­ern Cal­i­for­nia cetaceans. The Jour­nal of Par­a­sitol­ogy, 64/4: 593-596.

Fer­rero, R., W. Walker. 1996. Age, growth, and re­pro­duc­tive pat­terns of the Pa­cific white-sided dol­phin (Lagenorhynchus obliq­uidens) taken in high seas drift nets in the cen­tral North Pa­cific Ocean. Cana­dian Jour­nal of Zo­ol­ogy, 74/9: 1673-1687.

Gib­bons, E., B. Dur­rant, J. De­marest. 1995. Con­ser­va­tion of En­dan­gered Species in Cap­tiv­ity: An In­ter­dis­ci­pli­nary Ap­proach. New York: SUNY Press.

Ham­mond, P., G. Bearzi, A. Bjørge, K. For­ney, L. Karkz­marski, T. Ka­suya, W. Per­rin, M. Scott, J. Wang, R. Wells, B. Wil­son. 2012. "Lagenorhynchus obliq­uidens" (On-line). The IUCN Red List of Threat­ened Species. Ac­cessed March 27, 2015 at http://​www.​iucnredlist.​org/​details/​11145/​0.

Hayssen, V., A. Tien­hoven. 1993. As­dell's Pat­terns of Mam­malian Re­pro­duc­tion: A Com­pendium of Species-spe­cific Data. New York: Cor­nell Uni­ver­sity Press.

Hen­der­son, E. 2010. Cetaceans in the South­ern Cal­i­for­nia Bight: Be­hav­ioral, Acoustic, and Spa­tio-tem­po­ral Mod­el­ing (The­sis). San Diego, CA: Uni­ver­sity of Cal­i­for­nia.

Hobbs, R., L. Jones. 1993. Im­pacts of high seas drift­net fish­eries on ma­rine mam­mal pop­u­la­tions in the North Pa­cific. In­ter­na­tional North Pa­cific Fish­eries Com­mis­sion Bul­letin, 53/3: 409-434.

Mann, J. 2000. Cetacean So­ci­eties: Field Stud­ies of Dol­phins and Whales. Chicago, Illi­nois: Uni­ver­sity of Chicago Press.

Marino, L. 1999. Brain growth in the har­bor por­poise and Pa­cific white-sided dol­phin. Jour­nal of Mam­mal­ogy, 80/4: 1353-1360.

Mar­tin, W., C. Haun, H. Bar­rows, H. Cravioto. 1970. Ne­ma­tode dam­age to brain of striped dol­phin, Lagenorhynchus obliq­uidens. Trans­ac­tions of the Amer­i­can Mi­cro­scop­i­cal So­ci­ety, 89/2: 200-205.

Nowak, R. 2003. Walker's Ma­rine Mam­mals of the World. Mary­land: The Johns Hop­kins Uni­ver­sity Press.

Pre­vost, J. 1996. White-Sided Dol­phins. Min­nesota: ABDO Pub­lish­ing Com­pany.

Rech­steiner, E., D. Rosen, A. Trites. 2013. En­ergy re­quire­ments of Pa­cific white-sided dol­phins (Lagenorhynchus obliq­uidens) as pre­dicted by a bioen­er­getic model. Jour­nal of Mam­mal­ogy, 94/4: 820-832.

Ridg­way, S., R. Har­ri­son. 1998. Hand­book of Ma­rine Mam­mals: The Sec­ond Book of Dol­phins and the Por­poises. Am­s­ter­dam, the Nether­lands: El­se­vier BV.

Sal­vadeo, C., D. Lluch-Belda, A. Gomez-Gal­lardo, J. Ur­ban-Ramirez, C. MacLeod. 2010. Cli­mate change and a pole­ward shift in the dis­tri­b­u­tion of the Pa­cific white-sided dol­phin in the north­east­ern Pa­cific. En­dan­gered Species Re­search, 11/1: 13-19.

Stew­art, B., P. Clapham, J. Pow­ell, R. Reeves. 2002. Guide to Ma­rine Mam­mals of the World. New York: Knopf Dou­ble­day Pub­lish­ing.

Weigl, R. 2005. Longevity of Mam­mals in Cap­tiv­ity: from the Liv­ing Col­lec­tions of the World: A List of Mam­malian Longevity in Cap­tiv­ity. Stuttgart, Ger­many: E. Schweizer­bart'sche.

Wil­son, D., S. Ruff. 1999. The Smith­son­ian Book of North Amer­i­can Mam­mals. Canada: UBC Press.

Yeater, D., H. Hill, N. Baus, H. Far­nell, S. Kuczaj. 2014. Vi­sual lat­er­al­ity in bel­u­gas (Del­phi­napterus leu­cas) and Pa­cific white-sided dol­phins (Lagenorhynchus obliq­uidens) when view­ing fa­mil­iar and un­fa­mil­iar hu­mans. An­i­mal Cog­ni­tion, 17/6: 1245-1259.