Geographic Range
The native range of
Ameiurus melas
extends west from the Appalachian mountain range to Arizona, north to southern Canada,
and as far north and east as New York. It can be found as far south as northern Mexico,
and the distribution excludes all but the panhandle of Florida. Introduced populations
occur in parts of California and Nevada. It also has been introduced to parts of England.
- Biogeographic Regions
- nearctic
- palearctic
Habitat
Black bullheads occupy most freshwater habitats, from small farm ponds to large lakes.
They can inhabit many waters that are otherwise unsuitable for other fishes. They
can tolerate poorly oxygenated, polluted, turbid, and high temperature waters. Because
they are relatively small, black bullheads also occupy many small creeks and rivers.
They prefer soft bottoms (in creeks and rivers) and avoid free flowing waters where
water moves rapidly. They feed in waters from one to three meters deep.
- Habitat Regions
- temperate
- freshwater
- Aquatic Biomes
- benthic
- lakes and ponds
- rivers and streams
Physical Description
Black bullheads are distinguished by their robust, bulky head. They have a very broad
head with pigmented barbels. The color ranges from dark brown to black dorsally,
yellow to white ventrally. Color varies greatly depending on location and water features.
They have long barbels, which are completely pigmented, and nostril whiskers resembling
"horns". Their fins have black pigmentation, the caudal fin is rounded and occasionally
has a pale vertical stripe at its base. Anal fin rays range from 19 to 25, caudal
fin rays from 15 to 18. Gill rakers range from 16 to 18. Similar species include yellow
bullheads (
Ameiurus natalis
),and brown bullheads (
Ameiurus platycephalus
). Black bullheads are the only bullhead species (
Ameiurus
) with completely pigmented barbels. Yellow bullheads have no pigmentation and brown
bullheads have light pigments on the ends of the barbels. Black bullheads naturally
hybridize with brown bullheads.
- Other Physical Features
- ectothermic
- heterothermic
- bilateral symmetry
- venomous
- Sexual Dimorphism
- sexes alike
Development
After spawning, eggs hatch in 4 to 10 days. Free swimming fry remain close to the
adult male for around two weeks. During this time the young reach around 25 mm in
total length. Average growth is to 170 mm in the first year, 240 mm in the second
year, 290 mm in the third year, 320 mm in the fourth year, peaking at around 350 mm
total length by the fifth year. Sexual maturity is reached around 160 mm. Population
density greatly affects black bullheads and sizes may vary greatly due to this.
Reproduction
After a female has constructed a nest, she intices a male by nudging the male's abdomen
with her snout. After breeding, the pair lay side by side, with the male curling his
caudal fin around the females mouth. After several pairings, spawning can be noticed
by a quivering in the female. The female guards the nest for the first day, then the
male takes over for the remainder of egg and fry protection.
- Mating System
- monogamous
Spawning occurs between May and July. The female fans out a saucer shaped nest in
a soft substrate, then removes larger elements with her snout. The male is nearby
during the construction of the nest. Nests are typically in 2 to 4 feet of water and
range in diameter and depth according to the substrate. On few occasions nest lay
beneath a log or other forms of structure. Females produce between 2,000 and 3,800
eggs. Spawning occurs five times over a one hour period. The male watches over the
nest after the first day for up to ten days. When the eggs then hatch, they stay
close to the male for up to two weeks.
- Key Reproductive Features
- iteroparous
- seasonal breeding
- gonochoric/gonochoristic/dioecious (sexes separate)
- sexual
- fertilization
- oviparous
Prior to breeding, females construct a nest using pelvic and anal fins. After breeding
the female guards the nest for the first day. After the first day males take over
and guard the nest for up to 10 days until the eggs hatch. For the next two weeks
the young remain close to the male.
- Parental Investment
-
pre-fertilization
- provisioning
-
protecting
- male
- female
-
pre-hatching/birth
-
provisioning
- male
- female
-
protecting
- male
- female
-
provisioning
-
pre-independence
-
provisioning
- male
-
protecting
- male
-
provisioning
Lifespan/Longevity
Black bullheads have an average lifespan of five years in the wild and a slightly
higher lifespan in captivity. The oldest found are around ten years. They are easily
kept in aquariums and adapt well. If the proper space and living conditions are met,
many find the these fish thrive well in captivity.
Behavior
Adults are very inactive during daylight hours, feeding almost exclusively after dark,
and are seldom seen or caught in rivers and streams until after dusk. Blacks bullheads
tend to look for food after dark along with up to four others. No social systems
have been observed.
- Key Behaviors
- natatorial
- nocturnal
- crepuscular
- motile
- aestivation
Home Range
Little is known about the size of the home range in black bullheads. They tend to
utilize pools in rivers and occupy areas where food is available.
Communication and Perception
Black bullheads have taste buds on in the mouth that help differentiate prey items.
Barbels are used to pick up chemical and hydrodynamic cues left by prey. As in many
catfishes the swim bladder is used to pick up on vibrations, as well as communicate.
Food Habits
Young black bullheads usually thrive on
ostracods
,
amphipods
,
copepods
, and insects and their larva. Young feed primarly in schooling patterns during midday.
Adults tend to be nocturnal, and feed on a wide variety of invertebrates. Midge larvae
and other young insects are the primary diet for adult bullheads. Black bullheads
have been known to eat small fish and fish eggs as well.
- Primary Diet
-
carnivore
- insectivore
- eats non-insect arthropods
- Animal Foods
- fish
- eggs
- carrion
- insects
- terrestrial worms
- aquatic crustaceans
- Plant Foods
- macroalgae
- Other Foods
- detritus
Predation
Young black bullheads may fall prey to largemouth bass (
Micropterus salmoides
) and other
basses
, as well as walleye (
Stizostedion vitreum
). They are protected from some predation by their venomous pectoral spines, that
can inflict a painful sting.
Ecosystem Roles
Black bullheads raise turbidity levels in farm ponds. Because of this they can negatively affect other species which also inhabit the pond. Black bullheads can survive in muddy or turbid waters where many species do not thrive. Black bullheads are important intermediate predators in the ecosystems in which they live.
Economic Importance for Humans: Positive
Though black bullheads are relatively small, they have become a popular fish among
anglers. They are known for their good taste, and amount of fight when body size proportions
are considered. Many black bullheads are kept in captivity because they adapt well
and have a long lifespan.
Economic Importance for Humans: Negative
Black bullheads are not considered a problem to most humans. In some cases where
overpopulation is a problem, they may never reach acceptable angling size. Where stocked
in Europe, most populations are too dense to reach full size capacity, which makes
them generally an unpopular species. Black bullheads can cause a painful sting if
pectoral spines puncture human flesh. Black bullheads contain small amounts of venom
at the ends of spine which can cause pain for up to a week.
- Negative Impacts
- injures humans
Conservation Status
Black bullheads are common and sometime very abundant throught their range. They
have become a popular gamefish in many areas, so due to stocking in many ponds and
lakes black bullheads are a stable and growing species.
Additional Links
Contributors
Tanya Dewey (editor), Animal Diversity Web.
Chris Rose (author), Eastern Kentucky University, Sherry Harrel (editor, instructor), Eastern Kentucky University.
- Nearctic
-
living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.
- introduced
-
referring to animal species that have been transported to and established populations in regions outside of their natural range, usually through human action.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- Palearctic
-
living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.
- introduced
-
referring to animal species that have been transported to and established populations in regions outside of their natural range, usually through human action.
- temperate
-
that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).
- freshwater
-
mainly lives in water that is not salty.
- benthic
-
Referring to an animal that lives on or near the bottom of a body of water. Also an aquatic biome consisting of the ocean bottom below the pelagic and coastal zones. Bottom habitats in the very deepest oceans (below 9000 m) are sometimes referred to as the abyssal zone. see also oceanic vent.
- ectothermic
-
animals which must use heat acquired from the environment and behavioral adaptations to regulate body temperature
- heterothermic
-
having a body temperature that fluctuates with that of the immediate environment; having no mechanism or a poorly developed mechanism for regulating internal body temperature.
- bilateral symmetry
-
having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.
- venomous
-
an animal which has an organ capable of injecting a poisonous substance into a wound (for example, scorpions, jellyfish, and rattlesnakes).
- monogamous
-
Having one mate at a time.
- iteroparous
-
offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).
- seasonal breeding
-
breeding is confined to a particular season
- sexual
-
reproduction that includes combining the genetic contribution of two individuals, a male and a female
- fertilization
-
union of egg and spermatozoan
- external fertilization
-
fertilization takes place outside the female's body
- oviparous
-
reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.
- natatorial
-
specialized for swimming
- nocturnal
-
active during the night
- crepuscular
-
active at dawn and dusk
- motile
-
having the capacity to move from one place to another.
- visual
-
uses sight to communicate
- tactile
-
uses touch to communicate
- acoustic
-
uses sound to communicate
- chemical
-
uses smells or other chemicals to communicate
- visual
-
uses sight to communicate
- tactile
-
uses touch to communicate
- acoustic
-
uses sound to communicate
- chemical
-
uses smells or other chemicals to communicate
- carrion
-
flesh of dead animals.
- macroalgae
-
seaweed. Algae that are large and photosynthetic.
- detritus
-
particles of organic material from dead and decomposing organisms. Detritus is the result of the activity of decomposers (organisms that decompose organic material).
- pet trade
-
the business of buying and selling animals for people to keep in their homes as pets.
- food
-
A substance that provides both nutrients and energy to a living thing.
- venomous
-
an animal which has an organ capable of injecting a poisonous substance into a wound (for example, scorpions, jellyfish, and rattlesnakes).
- carnivore
-
an animal that mainly eats meat
- insectivore
-
An animal that eats mainly insects or spiders.
References
Etnier, D., W. Starnes. 1993. The Fishes of Tennessee . The University of Tennessee Press/ Knoxville.
2004. "A Boundary Waters Compendium" (On-line). Accessed October 31, 2005 at http://www.rook.org/earl/bwca/nature/fish/ictalurusmel.html .