Geographic Range
Corvus frugilegus
is found in the Palearctic region, across much of Europe and Asia. Two subspecies
of rooks are recognized:
C. f. frugilegus
and
C. f. pastinator
. The geographic range of
C. f. frugilegus
extends from Ireland eastward across Europe into Russia, with southern boundaries
as far as Turkey and Iran. Sporadic, localized populations can be found as far north
as southern Sweden, Norway, and Denmark. Populations have also been introduced in
New Zealand, where they flourish. During breeding and migration,
C. f. frugilegus
can be found in northern Russia and the Mediterranean area, respectively. The geographic
range of
C. f. pastinator
, commonly known as the Oriental rook, extends from eastern Asia west into northern
Mongolia. The two subspecies are generally geographically separated by the Altai Mountains.
- Biogeographic Regions
- palearctic
- oceanic islands
Habitat
Rooks are widely distributed across Europe and western Asia, preferring arable land,
river plains, and steppe regions where soil is generally soft and fertile. In agricultural
landscapes, rooks tend to avoid areas where winter cereal grains such as rye and wheat
are grown, instead preferring areas in which softer and more easily-accessible spring
cereals such as barley are grown. Spring cereals are an ideal food source for
C. frugilegus
due to easy foraging that results from their small height. River plains and steppe
regions also serve as excellent habitats for rooks because their rich soil is usually
teeming with insects, and soft ground makes foraging possible. Rooks can also be found
in areas bordering cities and towns as long as large trees are available for cover
and food is available for scavenging. Given their wide occupation of much of Europe
and Asia, rooks are able to tolerate a large elevation range, from sea level to approximately
4000 m.
- Habitat Regions
- temperate
- terrestrial
- Terrestrial Biomes
- savanna or grassland
- forest
- Other Habitat Features
- suburban
- agricultural
- riparian
Physical Description
Corvus frugilegus
is similar in appearance to
C. corone
(carrion crow), another species of
corvid
. Though both species are covered in glossy black feathers with a metallic sheen,
rooks are distinguishable from carrion crows by their slightly smaller size, distinct
wedge-shape tail, light colored bill, and prominently-fingered wingtips. Rooks average
47 cm long and weigh 337 to 531 g, but are considered large when compared to most
other corvid species. Rooks show weak sexual dimorphism, with males slightly larger
than females. In rooks, wing length ranges from 290 to 330 mm (wingspan ranges from
81 to 94 cm) and tarsus length ranges from 52 to 58 mm. For their size, rooks have
a relatively large bill (53 to 57 mm long) that tapers to a sharp point. This long,
sharp bill aids in food retrieval and eating insects.
Corvus frugilegus frugilegus
tends to have a longer, thicker beak than that of
C. f. pastinator
, as well as a larger area of bare skin covering the forehead, lores (skin between
eye and bill on side of head), and gular area (skin that joins lower mandible to neck).
Corvus frugilegus frugilegus
also has a violet sheen to its black feathers, whereas
C. f. pastinator
has a greenish sheen. Juveniles of both subspecies are easily recognizable by their
brown-toned feathers and fully-feathered face, which does not become bare until their
first spring.
- Other Physical Features
- endothermic
- homoiothermic
- bilateral symmetry
- Sexual Dimorphism
- sexes alike
- male larger
Reproduction
Despite their highly social nature, rooks form pair bonds that lasts from several
years to life. Rooks generally take mates when they are two years old. During the
fall mating season, pair bonds nest together in communal roosts called rookeries until
they return to individual nests to lay eggs. Despite hundreds of birds in a single
rookery, rooks maintain their pair bonds through extensive communication. Though rooks
are known for being monogamous, like other corvid species such as
Corvus corax
(common ravens) and
C. corone
(carrion crows), there have been reported instances of bigamy and occupation of a
nest by multiple females.
- Mating System
- monogamous
- cooperative breeder
Breeding and egg-laying usually begins around late February in Britain, but may be
as late as April and May in central Europe and Russia where cold weather persists
for a longer period of time. Rooks generally build nests in tall deciduous trees,
though nests on the ground and in bushes are not uncommon. Nests consist of sticks
and branches with a deep leaf, grass, and moss-lined cup. Rooks (
Corvus frugilegus
) lay two to seven (average four) blue-green eggs that are covered with brown and
grey mottling. Rook eggs are very similar in appearance to those of ravens (
C. corax
), though slightly smaller, on average 40 mm long. After 16 to 18 days of incubation
mainly by the female, the young hatch blind and helpless.
- Key Reproductive Features
- iteroparous
- seasonal breeding
- gonochoric/gonochoristic/dioecious (sexes separate)
- sexual
- oviparous
For the 16 to 18 day incubation period the female rook covers the eggs unless she
has to briefly leave the nest, in which case the male takes over this duty. After
hatching, the female tends to the young exclusively while the male delivers food.
This continues for approximately the first ten days until the young become more self-sufficient,
at which point the female joins the male in food gathering. At around 32 to 33 days
old the young rooks fledge and leave the nest, but roost in nearby trees to remain
close to the parents. The young continue their relationship with the parents for several
weeks until they become fully independent. Even after reaching full maturity and independence,
rooks generally remain members of their original rookery.
- Parental Investment
- altricial
- male parental care
- female parental care
-
pre-fertilization
- provisioning
-
protecting
- female
-
pre-hatching/birth
-
provisioning
- male
- female
-
protecting
- male
- female
-
provisioning
-
pre-weaning/fledging
-
provisioning
- male
-
protecting
- male
- female
-
provisioning
-
pre-independence
-
provisioning
- male
- female
-
protecting
- male
- female
-
provisioning
- extended period of juvenile learning
Lifespan/Longevity
Not much is recorded on the lifespan of
C. frugilegus
, but like most
Corvidae
species, the rook is expected to live 15 to 20 years in the wild. According to the
EURING European Longevity Records, the oldest rook found in the wild lived to be 22
years old. As is the trend, rooks in captivity may live for much longer. In a similar
species,
Corvus corax
, the longest lifespan was recorded at 69 years for life in captivity.
Behavior
Rooks are highly social, living and interacting in large groups, although mating tends to monogamous. This bird species is largely arboreal and actively defends its territory. Rooks are active primarily during the day.
Communication and Perception
Rooks have a distinct call that has been described as sounding like a “caw." When
rooks are defending or establishing a territory, multiple “caws” are used. Rooks also
have a snarling call, as well as a gull call, which are used when an intruder comes
within a short distance of their nest. In order to remain in contact with other rooks,
a single loud “caw” is used during both foraging and migration. However, it is believed
that rooks mainly vocalize with their mate, rather than in other social interactions.
Like other birds, rooks perform singing duets (usually with a mate) that are believed
to create a stronger mating bond. Female rooks use frequent vocalizations, by means
of a begging call, in order to establish a submissive state with their mate, as well
as to show dependency towards the male. Rooks also rely heavily on vocal communication
with their young during the first few days of hatching.
Auditory communication is vital to the rook. They have a sense of hearing which helps
them distinguish amongst other populations and species. The rook is able to recognize
the call of a mate or its young. In addition to vocalizations, rooks also rely on
visual communication, which becomes increasingly more important once young are able
to open their eyes. Pecking is another form of communication, when an intruder rook
comes too close to a territory; pecking attacks can occur usually resulting in the
retreat of the intruder.
- Other Communication Modes
- duets
- Perception Channels
- visual
- polarized light
- tactile
- acoustic
- chemical
Food Habits
Rooks are opportunistic feeders. As omnivores, they eat any edible food item. Due
to the strength and size of the bill, rooks are often found probing the ground in
search of earthworms or other insects. Rooks also ingest small acorns, small fruits,
and cereal grains. When the opportunity arises, rooks prey on small mammals, small
birds, carrion, and eggs of the same species. They also have been known to act as
“nest predators”, attacking the nests of other species of birds in order to eat the
hatchlings and eggs.
Rook feeding habits often vary due to the location of their nest. Unlike those occurring
in natural areas as above, those that live near urban sites also act as scavengers
and take advantage of trashcans as well as abandoned food. Most rooks spend much of
their time foraging at dawn and dusk. Primarily searching at dawn, rooks will pick
through garbage bags to obtain food. However, they have been seen foraging during
the day. Like all
corvids
, rooks store their food.
- Primary Diet
-
carnivore
- eats terrestrial vertebrates
- eats eggs
- insectivore
- eats non-insect arthropods
- vermivore
- scavenger
- herbivore
- omnivore
- Animal Foods
- birds
- mammals
- eggs
- carrion
- insects
- terrestrial worms
- aquatic crustaceans
- Plant Foods
- seeds, grains, and nuts
- fruit
- Foraging Behavior
- stores or caches food
Predation
Little is known about predators of rooks. However, they appear to have similar predators
to
Corvus corax
or
Corvus brachyrhynchos
, both similar and closely related species in the Family
Corvidae
. Owls, such as
Bubo virginianus
, hawks, or even intruding species of
Corvidae
tend to be the primary predators of rooks. Raptorial bird species prey on fledglings
from nests more often than attacking adults. Humans may also pose as a threat to some
species due to increasing tolerance of human presence. Humans are a threat due to
shootings and habitat destruction of rooks.
Ecosystem Roles
Rooks have numerous roles in the ecosystem. They serve as hosts for numerous protozoan
organisms such as trypanosomes and leucocytozoans. Such organisms are generally not
pathogenic and merely occupy rooks as vectors. While rook hosting of these organisms
does not directly cause it any harm, it makes infection of other species possible.
In this way, rooks sustain the lifecycle of these organisms.
Rooks also serve as hosts for oribatid mites, among other types of mites. Oribatid
mites are soil mites, and feed on dead plant and fungal material. These mites live
in the feathers of rooks, where it is believed they consume fungi. Such a relationship
does not harm rooks, it is actually beneficial, though its results are not readily
noticeable.
Lastly, rooks play a key role in seed dispersal as they crack open and consume cereal
grains such as barley. Without birds and small animals to crack open seeds and remove
them from the plant, seed dispersal for cereal grains could be problematic or less
efficient.
- Ecosystem Impact
- disperses seeds
- soil aeration
- oribatid mites ( Oribatidae )
- trypanosomes ( Trypanosoma )
- leucocytozoans ( Leucocytozoon )
Economic Importance for Humans: Positive
While many farmers claim that
C. frugilegus
does more harm than good, recent studies suggest that 60 to 90% of insects consumed
by rooks are agricultural pests. If this is the case, large numbers of rooks may have
some impact on pest insect populations. Rooks are also known to dig into the soil
in search of insects, so this may have a slight aeration effect which is particularly
important in agricultural environments. Lastly, rooks play an important role in seed
dispersal as they consume and crack open cereal grains.
- Positive Impacts
- controls pest population
Economic Importance for Humans: Negative
Rooks are commonly referred to as "agricultural pests", meaning they cause the loss
and destruction of commercial crops. When foraging for food, rooks are often found
in farmland crops, taking advantage of the cereals and grains. This can lead to an
economic decline for farmers, as well as any person or company that may use the farmer
for food. However, rooks are only known to cause damage to crops if the preferred
food is not available.
As well as being agricultural pests, rooks that live in urban areas are likely to
get into garbage and rip open the bags, which can in turn cause problems for humans.
- Negative Impacts
- crop pest
Conservation Status
Corvus frugilegus
is classified as a species of least concern on the IUCN's Red List of Threatened
Species.
Corvus frugilegus
is abundant and is able to maintain stable populations in its habitat.
Additional Links
Contributors
Lauren Carlson (author), Radford University, Kelsey Townsend (author), Radford University, Christine Small (editor), Radford University, Tanya Dewey (editor), University of Michigan-Ann Arbor.
- Palearctic
-
living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- oceanic islands
-
islands that are not part of continental shelf areas, they are not, and have never been, connected to a continental land mass, most typically these are volcanic islands.
- introduced
-
referring to animal species that have been transported to and established populations in regions outside of their natural range, usually through human action.
- temperate
-
that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).
- terrestrial
-
Living on the ground.
- tropical savanna and grassland
-
A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.
- savanna
-
A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.
- temperate grassland
-
A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.
- forest
-
forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.
- suburban
-
living in residential areas on the outskirts of large cities or towns.
- agricultural
-
living in landscapes dominated by human agriculture.
- riparian
-
Referring to something living or located adjacent to a waterbody (usually, but not always, a river or stream).
- endothermic
-
animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.
- bilateral symmetry
-
having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.
- monogamous
-
Having one mate at a time.
- cooperative breeder
-
helpers provide assistance in raising young that are not their own
- iteroparous
-
offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).
- seasonal breeding
-
breeding is confined to a particular season
- sexual
-
reproduction that includes combining the genetic contribution of two individuals, a male and a female
- oviparous
-
reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.
- altricial
-
young are born in a relatively underdeveloped state; they are unable to feed or care for themselves or locomote independently for a period of time after birth/hatching. In birds, naked and helpless after hatching.
- male parental care
-
parental care is carried out by males
- female parental care
-
parental care is carried out by females
- arboreal
-
Referring to an animal that lives in trees; tree-climbing.
- diurnal
-
- active during the day, 2. lasting for one day.
- motile
-
having the capacity to move from one place to another.
- migratory
-
makes seasonal movements between breeding and wintering grounds
- territorial
-
defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement
- social
-
associates with others of its species; forms social groups.
- visual
-
uses sight to communicate
- tactile
-
uses touch to communicate
- acoustic
-
uses sound to communicate
- duets
-
to jointly display, usually with sounds in a highly coordinated fashion, at the same time as one other individual of the same species, often a mate
- visual
-
uses sight to communicate
- polarized light
-
light waves that are oriented in particular direction. For example, light reflected off of water has waves vibrating horizontally. Some animals, such as bees, can detect which way light is polarized and use that information. People cannot, unless they use special equipment.
- tactile
-
uses touch to communicate
- acoustic
-
uses sound to communicate
- chemical
-
uses smells or other chemicals to communicate
- carrion
-
flesh of dead animals.
- stores or caches food
-
places a food item in a special place to be eaten later. Also called "hoarding"
- soil aeration
-
digs and breaks up soil so air and water can get in
- carnivore
-
an animal that mainly eats meat
- insectivore
-
An animal that eats mainly insects or spiders.
- scavenger
-
an animal that mainly eats dead animals
- herbivore
-
An animal that eats mainly plants or parts of plants.
- frugivore
-
an animal that mainly eats fruit
- granivore
-
an animal that mainly eats seeds
- omnivore
-
an animal that mainly eats all kinds of things, including plants and animals
References
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