Geographic Range
Eleonora’s falcons are distributed throughout the Mediterranean region, including
the Canary Islands, Morocco, Algeria, Tunisia, the Balearic Islands, Sicily, Sardinia,
Croatia, islands in the Aegean Sea, Crete, Cyprus, and Turkey. All breeding sites
lie between longitude 14°W to 33°E, and latitude 43°N to 28°N. Approximately 10% of
Eleonora falcon populations breed on the Tilos Island chain. All populations winter
in Madagascar and the Mascarene Islands.
- Biogeographic Regions
- palearctic
- ethiopian
- mediterranean sea
Habitat
Eleonora’s falcons commonly breed on the ground on small, rocky, undisturbed islands
with little vegetation. On larger islands, such as Sicily and Sardinia, and on the
mainland in Africa and Turkey, they seek inaccessible coastal cliffs to breed. They
hunt over the open sea, as well as over wetlands, grasslands, and sometimes woodlands.
Nest sites are found from sea-level to an elevation of 2,000 meters.
- Habitat Regions
- temperate
- tropical
- terrestrial
- Terrestrial Biomes
- savanna or grassland
- forest
- Aquatic Biomes
- coastal
Physical Description
Eleonora’s falcons are medium-sized falcons, of similar length to peregrine falcons
(
F. peregrinus
), but much lighter in weight and build. Like all other falcons, Eleonora’s falcons
show reversed sexual size dimorphism, with females on average slightly larger than
males. Males vary in length from 37 to 43 cm, females from 38.5 to 45.5 cm, and in
weight, males range from 350 to 390 g, females from 340 to 460 g. The shortest wingspan
measurement for an Eleonora’s falcon was 84 cm, while large females may have a wingspan
of up to 103 cm. Eleonora’s falcons have long, narrow wings that exceed the tip of
the long tail when folded. There are two color morphs, a light and a dark morph. Dark
morph birds are all brown, and may appear black from a distance. The male’s cere is
yellow in color while the female’s is blue-gray. Light morph birds are also dark brown
above and buff to rufous below with dark streaking. The light morph also shows the
typical dark moustache stripe.
- Other Physical Features
- endothermic
- homoiothermic
- bilateral symmetry
- polymorphic
- Sexual Dimorphism
- female larger
Reproduction
Eleonora’s falcons are monogamous, colonial breeders. Aerial displays by male falcons
begin as soon as the birds arrive on nesting sites.
- Mating System
- monogamous
Breeding sites are occupied starting in late April, though breeding does not start
until late July. The young hatch in late August to early September at the beginning
of the migration period of most small, Palearctic birds. The nest is located on the
ground or on a cliff, often in a small cavity or under a small bush, sheltered from
wind. Clutch size ranges from 1 to 4 eggs, rarely 5. Incubation lasts between 28 and
30 days, and the young fledge after another 35 to 40 days. Average productivity differs
among different colonies and can range from 1.26 fledglings per year to 2.6 young
per year. Young males usually remain near their parents’ home range, while females
disperse farther.
- Key Reproductive Features
- iteroparous
- seasonal breeding
- gonochoric/gonochoristic/dioecious (sexes separate)
- sexual
- oviparous
As in most other
falcon
species, both males and females contribute to incubating, protecting, and nourishing
their offspring.
- Parental Investment
- altricial
-
pre-fertilization
- provisioning
-
protecting
- male
- female
-
pre-hatching/birth
-
provisioning
- male
-
protecting
- male
- female
-
provisioning
-
pre-weaning/fledging
-
provisioning
- male
-
protecting
- male
- female
-
provisioning
-
pre-independence
-
provisioning
- male
- female
-
provisioning
Lifespan/Longevity
There is no information available on lifespan in Falco eleonorae .
Behavior
Eleonora’s falcons are gregarious. They nest in colonies of up to 300 pairs and may
hunt migrating passerines and invertebrates singly or in small groups. Members of
the colony aggressively chase off other birds, such as doves and gulls. When not occupying
breeding colonies, these falcons are likely to be solitary.
Migration starts late for Eleonora’s falcons, they usually leave the breeding grounds
in late October to early November and migrate to Madagascar and the Mascarene Islands.
Little is known about their migration route, although it is likely that they follow
the east African coastline.
Home Range
The breeding territories of Eleonora’s Falcons vary within each colony. Some territories
are as small as 1 m² while others are up to 200 m². This space is defended by the
local pair against any intruders, including other falcons. The air space up to 10
m in height above this area is also protected by the resident pair. The nesting cliff
is defended by all members of the colony, and hunting falcons may fly up to 5 km from
the colony to chase passerines. Individual birds have been observed to travel up to
30 km from the breeding cliff to hunt.
Communication and Perception
Eleonora’s falcons are highly vocal and seem to have a wider range of vocalizations
than other falcons. Vocal communication is especially common during the pre-mating
season. Some common calls include recognition, territorial, display, copulation, greeting,
prey transfer demand, prey arrival, alarm, and distress calls. Chuckles can also be
heard by brooding falcons upon arrival of the mate. The young give a variety of calls
to indicate discomfort, hunger, alarm, or excitement.
Eleonora's falcons, like other falcons, have especially keen vision, used in capturing prey.
Food Habits
Eleonora’s falcons feed primarily on flying insects. During the breeding season, however,
these falcons switch their hunting behavior and concentrate solely on migrating passerine
birds. Over 100 species have been recorded as prey of Eleonora’s falcons. To hunt,
falcons fly out to sea utilizing vertical winds for soaring, and may hunt close to
sea level or up to a height of 1,000 m. Several falcons often hunt together spaced
several hundred meters apart, creating a barrier which makes it harder for migrating
birds to stay undetected. Eleonora’s falcons are also known to hunt until well after
sunset. These falcons usually stoop down on their prey, but sometimes pursue it for
longer distances. Insects are also caught and eaten in flight.
During the breeding season excess prey is often cached and retrieved later. Male falcons
do most of the bird-hunting during the breeding season, leaving females to incubate
eggs and nestlings.
- Primary Diet
-
carnivore
- eats terrestrial vertebrates
- insectivore
- Animal Foods
- birds
- insects
- Foraging Behavior
- stores or caches food
Predation
Eleonora’s falcons have few natural predators as adults. Black rats (
Rattus rattus
) inhabit many islands that harbor falcon colonies and may feed on unguarded falcon
eggs or nestlings. Peregrine falcons (
Falco peregrinus
), Lanner falcons (
Falco biarmicus
), and Eurasian eagle owls (
Bubo bubo
) have been known to prey on adult falcons. Cory’s shearwaters (
Calonectris diomedea
) often breed within falcon colonies and occasionally take young falcons.
Ecosystem Roles
Eleonora’s falcons play an important role in the migration of millions of small birds,
mainly
passerines
from their Palearctic breeding grounds to African wintering areas. Falcons prey on
migrating passerines, preferring small, juvenile birds. It is estimated that all Eleonora’s
falcon colonies combined catch about two million migrating birds in a single breeding
season. This, however, only represents 0.02 to 0.04% of the total number of migrating
birds.
Falco eleonorae
is also important in regulating prey species populations throughout their range.
Economic Importance for Humans: Positive
Eleonora’s falcons, like other raptors, are important bioindicators of healthy environments.
They further help to reduce pest species, such as
grasshoppers
and
rodents
that cause damage to human crops.
- Positive Impacts
- research and education
- controls pest population
Economic Importance for Humans: Negative
There are no negative impacts of Eleonara's falcons on humans.
Conservation Status
Due to the limited distribution and colonial breeding habits of Eleonora’s falcons,
they are very vulnerable. The main threats are an increase in tourism, especially
the use of motorboats near colonies, which often results in distressed parent birds
and reproductive failure. In the Aegean Sea, colonies are often raided by fisherman
who take the young and eggs for food. Persecution of adult birds has also been recorded.
The total population is currently estimated at 4,500 breeding pairs. As long as breeding
colonies are protected from poaching by humans and from heavy tourism, Eleonora’s
falcon populations will likely remain stable. A decrease in migrating passerines due
to habitat loss in mainland Europe has been observed over the past century and falcon
populations have dropped accordingly, but seem to be currently stable.
Additional Links
Contributors
Tanya Dewey (editor), Animal Diversity Web.
Dennis Lorenz (author), Michigan State University, Pamela Rasmussen (editor, instructor), Michigan State University.
- Palearctic
-
living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- Ethiopian
-
living in sub-Saharan Africa (south of 30 degrees north) and Madagascar.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- temperate
-
that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).
- tropical
-
the region of the earth that surrounds the equator, from 23.5 degrees north to 23.5 degrees south.
- terrestrial
-
Living on the ground.
- tropical savanna and grassland
-
A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.
- savanna
-
A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.
- temperate grassland
-
A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.
- forest
-
forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.
- coastal
-
the nearshore aquatic habitats near a coast, or shoreline.
- marsh
-
marshes are wetland areas often dominated by grasses and reeds.
- swamp
-
a wetland area that may be permanently or intermittently covered in water, often dominated by woody vegetation.
- endothermic
-
animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.
- bilateral symmetry
-
having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.
- polymorphic
-
"many forms." A species is polymorphic if its individuals can be divided into two or more easily recognized groups, based on structure, color, or other similar characteristics. The term only applies when the distinct groups can be found in the same area; graded or clinal variation throughout the range of a species (e.g. a north-to-south decrease in size) is not polymorphism. Polymorphic characteristics may be inherited because the differences have a genetic basis, or they may be the result of environmental influences. We do not consider sexual differences (i.e. sexual dimorphism), seasonal changes (e.g. change in fur color), or age-related changes to be polymorphic. Polymorphism in a local population can be an adaptation to prevent density-dependent predation, where predators preferentially prey on the most common morph.
- monogamous
-
Having one mate at a time.
- iteroparous
-
offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).
- seasonal breeding
-
breeding is confined to a particular season
- sexual
-
reproduction that includes combining the genetic contribution of two individuals, a male and a female
- oviparous
-
reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.
- altricial
-
young are born in a relatively underdeveloped state; they are unable to feed or care for themselves or locomote independently for a period of time after birth/hatching. In birds, naked and helpless after hatching.
- diurnal
-
- active during the day, 2. lasting for one day.
- crepuscular
-
active at dawn and dusk
- motile
-
having the capacity to move from one place to another.
- migratory
-
makes seasonal movements between breeding and wintering grounds
- solitary
-
lives alone
- colonial
-
used loosely to describe any group of organisms living together or in close proximity to each other - for example nesting shorebirds that live in large colonies. More specifically refers to a group of organisms in which members act as specialized subunits (a continuous, modular society) - as in clonal organisms.
- visual
-
uses sight to communicate
- acoustic
-
uses sound to communicate
- stores or caches food
-
places a food item in a special place to be eaten later. Also called "hoarding"
- carnivore
-
an animal that mainly eats meat
- insectivore
-
An animal that eats mainly insects or spiders.
- visual
-
uses sight to communicate
- tactile
-
uses touch to communicate
- acoustic
-
uses sound to communicate
- chemical
-
uses smells or other chemicals to communicate
References
Cade, T., A. Clark. 1979. Birds of Prey. Science , 206/4415: 211-212.
Ferguson-Lees, J., D. Christie. 2001. Raptors of the World . Boston, New York: Houghton Mifflin Company.
Hedenström, A., M. Rosen, S. Ǻkesson, F. Spina. 1999. Flight Performance During Hunting Excursions in Eleonora’s Falcon (Falco eleonorae). The Journal of Experimental Biology , 202: 2029-2039.
Hedenström, A., M. Rosen. 2001. Predator versus prey: On aerial hunting and escape strategies in birds. Behavioral Ecology , 12/2: 150-156.
Martinez, A., D. Oro, V. Ferris, R. Belenguer. 2002. Is growing tourist activity affecting the distribution or number of breeding pairs in a small colony of the Eleonora’s Falcon?. Animal Biodiversity and Conservation , 25/2: 47-51.
Massa, B. 1978. Observations on Eleonora’s Falcon (Falco eleonorae) in Sicily, Italy and Surrounding Islets. Ibis , 120/4: 531-534.
Mentzelopoulos, K. 2006. "Tilos Park" (On-line). Accessed January 08, 2007 at http://www.tilos-park.org/about.htm .
Palacios, C. 2004. Current status and distribution of birds of prey in the Canary Islands. Bird Conservation International , 14/3: 203-213.
Piasevoli, G., V. Scetaric. 2001. Eleonora’s Falcon (Falco eleonorae) in Croatia: Range, threats and the proposal of action and management plan. Annales Series Historia Naturalis , 23: 81-86.
Rohwer, S., D. Paulson. 1987. The Avoidance-Image Hypothesis and Color Polymorphism in Buteo Hawks. Ornis Scandinavica , 18/4: 285-290.
Rosen, M., A. Hedenström. 2002. Soaring Flight in the Eleonora’s Falcon (Falco eleonorae). The Auk , 119/3: 835-840.
Rosen, M., et al. 1999. Hunting flight behavior of the Eleonora’s Falcon (Falco eleonorae). Journal of Avian Biology , 30/4: 342-350.
Swatscheck, I., et al. 1993. Population Genetics and Paternity Analysis of Eleonora’s Falcon (Falco eleonorae). Journal fuer Ornithologie , 134/2: 137-143.
Thiollay, J., B. Meyburg. 1981. Organization of an Island Population of Raptors in Madagascar. Alauda , 49/3: 216-226.
Walter, H. 1979. Eleonora's Falcon . Chicago: The University of Chicago Press.
Wink, M., C. Wink, D. Ristow. 1982. Biology of the Eleonora’s Falcon (Falco eleonorae): Biometrics of Sexual Dimorphism of Adult and Fledged Falcons. Vogelwelt , 103/6: 225-229.
Wink, M., C. Wink, D. Ristow. 1982. Biology of the Eleonora’s Falcon (Falco eleonorae): Breeding Success in Relation to Nest Site Exposition. Journal fuer Ornithologie , 123/4: 401-408.
Wink, M., C. Wink, D. Ristow. 1980. Biology of the Eleonora’s Falcon (Falco eleonorae): Clutch Size in Relation to Hunting Success and Weight of the Parent Falcons. Journal fuer Ornithologie , 121/4: 387-390.