Geographic Range
Buffleheads are native to North America. Their summer breeding range includes central
Alaska and extends south to British Columbia and east to Saskatchewan. Isolated breeding
populations can also be found throughout the northern United States and in Quebec.
Their winter distribution is generally split into two populations, one on the east
coast and the other on the west coast of North America. The east coast population
is usually found from New Jersey to North Carolina, and can be reaches up to the Bay
of Fundy in New Brunswick, Canada. The west coast population is concentrated in British
Columbia, Washington, and California. The most substantial winter confluence occurs
on British Columbia’s Vancouver Island and the Californian coast. They are less likely
to be found moving inland from the western Klamath Basin in California and Oregon
toward the Mississippi River.
Habitat
Buffleheads live in boreal forests and aspen parklands as well as seasonally-flooded
wetlands and estuaries. They can be found along ecotones, and in marshes, farmlands,
grasslands, and open waters. They prefer ponds and small lakes with no drainage while
breeding. During migration, they use rivers and available water bodies as temporary
habitat. Their winter habitat includes salty bodies of water like marshes, coastlines,
and estuaries with shelter. They are not found at high mountain elevations.
- Habitat Regions
- temperate
- terrestrial
- saltwater or marine
- freshwater
- Terrestrial Biomes
- taiga
- savanna or grassland
- forest
- scrub forest
- Aquatic Biomes
- lakes and ponds
- rivers and streams
- temporary pools
- coastal
- brackish water
- Other Habitat Features
- suburban
- agricultural
- riparian
- estuarine
Physical Description
Buffleheads are small diving ducks that exhibit strong sexual dimorphism. Males with
breeding plumage are predominately black and white, with a black head and back that
appears iridescent green and purple. They have a white underbelly and a distinguishing
large white patch extending from the nape of the neck to the crown of the head. Males
have blue-gray bills and pink webbed feet. Females are similar in plumage to male
yearlings. They are grey on the bottom and brown on top with a white patch on the
sides of the head. Both male yearlings and adult females have bills that are dark
gray to black and legs and toes that are dark pink while webbed feet are brown. The
ear patch of female buffleheads is more defined than the that of yearling males. The
downy coats of hatchlings are black to dark grey with a white patched cheeks, throats,
lower breasts, and bellies.
Buffleheads weigh 270 to 513 grams and are 32 to 40 cm long. Their wingspan is 16.9
to 17.5 cm long. Sexual dimorphism is exhibited in their size as well. Adult males
weigh 450 grams on average and are 35 to 40 cm long, while females weigh 325 grams
on average and are 32 to 35 cm long. Their folded wings are 18 cm or less in adults
and their tails are less than 8 cm long.
Adult males are sometimes mistaken for common goldeneyes (
Bucephala clangula
), Barrow’s goldeneyes (<< Bucephala islandica>>) and hooded mergansers (
Lophodytes cucullatus
). Unlike buffleheads, goldeneyes have a white patch that starts below the eye and
extends towards the beak and they both have golden eyes. Hooded mergansers are larger
and have a fan-shaped white patch on their heads. Unlike bufflehead males, their chests
and wings haave white stripes and a brownish or golden brown underside.
- Other Physical Features
- endothermic
- bilateral symmetry
- Sexual Dimorphism
- male larger
- sexes colored or patterned differently
- male more colorful
Reproduction
Buffleheads generally form a mating pair that stays together during the season and
in subsequent seasons. Less frequently, males pair with a second female after the
first has finished laying her eggs. Courtship behavior occurs throughout the year
and facilitates seasonal pairing of couples. Buffleheads use an array of physical
displays and vocalizations during courtship. Males bob their heads and fly low over
females to display their black and white underside and their pink legs, and then land
with their feet straight as if water skiing. Paired birds display a “following” behavior
where the female swims behind the male. The male stretches his neck upward and the
female extends her neck back while she follows behind him. Buffleheads use displays
both before and after copulation. Males also perform displays after threat or aggression
from other males.
- Mating System
- monogamous
Buffleheads breed once per year from late winter to early April. Breeding females
lay a single clutch between late April and mid-May with an average of 9 eggs per clutch.
Female buffleheads typically lay their eggs close to the same date each year, but
second and third-year breeding females lay 4 to 9 days earlier than first year breeders.
Their eggs are olive-buff colored. On average, they measure 50 by 36 mm and weigh
36.68 g. Females incubate the eggs for 30 days while males leave to molt. When leaving
the nest to feed during the incubation period, females cover their eggs with feathers.
Buffleheads use nests constructed by other species. Their nests are hollowed out cavities
in trees usually within 15 m of a body of water and above flood plain level. Nests
are often found in poplars and aspen, although pine trees are a favorite in the western
United States. Nests are bare and buffleheads do not add material to their nests.
Female buffleheads scout out their nest location up to a year in advance. If the desired
nest is occupied when she returns, she searches for a new site with the male. Females
only 1 year old have been observed searching for potential nest sites although they
do not begin breeding until age 2. Two females have been documented sharing a same
nest; however, one may evict the other that leaves the nest to feed. The average nest
entrance is approximately 7 cm in diameter and the cavity diameter is 11.5 to 21 cm
with a depth around 33.8 cm. Larger cavities are normally avoided because they are
favored by goldeneyes (
Bucephala islandica
and
Bucephala clangula
). Goldeneyes can kill buffleheads in larger nests, but cannot enter the smaller entrances
where buffleheads nest.
Bufflehead chicks hatch after 28 to 35 days. They typically hatch within a span of
12 hours, but may take up to 36 hours from first to last. It is thought that late
hatching eggs are laid during incubation. Precocial chicks are born with their eyes
open and fully covered in down, with a mass around 23.8 g. Buffleheads are able walk
as soon as their plumage dries out. Young buffleheads are nurtured intently during
and after hatching. Newly hatched chicks live in the nest for 1 to 2 days and are
then encouraged to jump from the nest hole. New mother buffleheads protect their
brood for 3 to 6 weeks, at which point the young buffleheads are considered independent.
Young ducklings are good swimmers, feeding on insects (92-100% of the time) on the
water and dabbling for vegetation. Their diving abilities develop in the first few
days and become the predominant mode of feeding. In fact, downy young spend 24% of
their time diving. Growth rates vary among individuals which is typical of precocial
birds. By day 20, the juvenile contour feathers begin to emerge with wing feathers
appearing at day 23. Their belly feathers appear next and the head, back and neck
feathers appear last. By day 40, males are apparently larger than their sisters. Plumage
is complete after day 50 and the chicks fledge in 45 to 55 days. They reach reproductive
maturity at 2 years.
- Key Reproductive Features
- iteroparous
- seasonal breeding
- gonochoric/gonochoristic/dioecious (sexes separate)
- sexual
- oviparous
Male buffleheads stay with their mates during egg laying and for part of the incubation
period. Females alone attend to the brood and defend their territory. They nurture
their young for the first 2 to 3 weeks after hatching. The young huddle tightly together
on both sides of the female on the shore or a floating log. Young buffleheads gather
tightly and close behind their mother if she gives an alarm call. In British Columbia,
34% of broods had at least one exchange of young buffleheads between mothers, usually
during a fight between them. Occasionally an entire brood is acquired by a female
bufflehead that won a territorial fight. Bufflehead mothers protect their brood for
up to 6 weeks, when the chicks are independent.
- Parental Investment
- precocial
- female parental care
-
pre-fertilization
-
protecting
- male
-
protecting
-
pre-hatching/birth
-
protecting
- male
- female
-
protecting
-
pre-weaning/fledging
-
provisioning
- female
-
protecting
- female
-
provisioning
-
pre-independence
-
provisioning
- female
-
protecting
- female
-
provisioning
Lifespan/Longevity
Buffleheads live an estimated 2.5 years for males and 2.3 years for females. In rare
cases, adults live very long and the record for the oldest adult is 18.7 years old.
However, information on survivorship is limited. The most recent survivorship data
available (1969 to 1973) was generated in New York State and was obtained by banding
and recovering birds. From these data it was estimated that annual survivorship of
females is between 61 to 73%, and 58 to 70% for males. Because the sample size was
particularly small, (56 to 159 birds of each sex) the accuracy of these estimates
is not known.
Behavior
Buffleheads spend their time preening, swimming, diving, perching, flying and foraging.
They seldom walk on land. They are moderately gregarious outside of the breeding season,
often found in flocks to 5 to 10 and rarely in flocks of more than 50. Their principal
daytime activity is foraging, comprising roughly 60 percent of their time. In the
winter, buffleheads continue to forage through the night. They also migrate at night
like many other ducks. Buffleheads have webbed feet that are used for propulsion during
swimming and diving, as well as treading water when startled or to gain speed for
flight. Nesting females perch at the opening of their nest cavities. Buffleheads preen
and bathe while on the surface of the water and adults without nests may sleep on
the surface of the water or on the shoreline. Females only sleep on land in bouts
of 20 to 30 minutes, particularly during egg-laying periods and while protecting their
young.
Males sometimes exhibit aggressive behavior and displays, which is explained by two
competing hypotheses. One suggests that territories are aggressively defended by male
buffleheads to secure a food resource or to protect the breeding ground. The other
hypothesis suggests that males are guarding the females, termed “mate-guarding.”
Males demonstrate a threatening head-forward position to protect nesting territory
sometimes accompanied by bumping and aggressive wing flapping. Bufflehead males may
also dive or fly directly at an opponent to protect their defined territory. Other
head postures such as an arching back or wing-flapping are considered a submissive
response by the loser of a territorial battle. Once females lay their eggs, males
do not defend as aggressively. After the hatchlings are out of the nest, males have
already left for molting grounds. Female then become the defenders of their broods.
Breeding territory of 0.33 hectares is most readily protected.
- Key Behaviors
- arboreal
- flies
- glides
- natatorial
- diurnal
- nocturnal
- crepuscular
- motile
- migratory
- territorial
- social
Home Range
Territory size in British Columbia averages .56 hectares, but only .38 hectares is
used with an intensely used territory of .38 ha. Males and females have different
territories. Females claim a territory in the first few days following hatching on
small ponds less than 400 m from the nest. This area is intensely defended by the
female against goldeneyes (
Bucephala clangula
and
Bucephala islandica
) and other ducks.
Communication and Perception
Buffleheads find prey underwater by sight, and they communicate through vocalizations
and displays. Courting bufflehead males bob their heads and produce a loud raspy noise.
In late winter and spring, they emit a low snarling grunts. Females make a loud deep
throated vocalization while following males during leading displays. Females use a
distinct low note to call their young which speeds up and increases in volume if she
becomes distressed. Buffleheads display a head-forward posture and raised wing feathers
when they are threatened or when protecting their territory or brood. Males protect
their territory by posturing next to other male buffleheads. This includes a head-forward
posture, flapping wings, and a raised tail. Once this display ends, they part ways.
Wing beating following the toe to toe display is thought to be a sign of concession
by the loser.
- Other Communication Modes
- duets
Food Habits
Buffleheads primarily eat aquatic invertebrates and some seeds. Their freshwater diet
consists of mostly insects like
damselfly larvae
,
dragonfly
larvae,
midge
larvae,
water boatmen
,
mayfly
larvae,
caddisfly
larvae and other insects. In saltwater habitats, a variety of
arthropods
and
molluscs
make up their diet. Buffleheads on the Pacific coast have been recorded consuming
herring
eggs in multi-species flocks and they occasionally eat fish like
sculpins
and
ratfish
. Prey is swallowed while submerged under the water. Buffleheads prefer to feed in
water less than 3 meters deep. All of their food is acquired by diving except for
downy young, who will dabble when first taking to the water.
Buffleheads' diet varies seasonally and by habitat. In the fall, pondweed seeds, sedges,
bulrushes, and mare’s tail become important to the bufflehead diet. The literature
reports that avian egg shells and bones have also been found in their stomachs. Female
buffleheads that consumed mostly gastropods during egg-laying were found to have higher
egg production. Eggs were also found to be larger with stronger shells. Gastropod
consumption peaks during incubation.
- Primary Diet
-
carnivore
- insectivore
- eats non-insect arthropods
- molluscivore
- Animal Foods
- amphibians
- fish
- insects
- mollusks
- aquatic crustaceans
- other marine invertebrates
- Plant Foods
- roots and tubers
- wood, bark, or stems
- seeds, grains, and nuts
Predation
Buffleheads are vulnerable to an assortment of predators that include birds of prey
and mammals. Included in this list are peregrine falcons (
Falco peregrinus
), snowy owls (
Nyctea scandiaca
), and bald eagles (
Haliaeetus leucocephalus
), and possibly great horned owls (
Bubo virginianus
) and Cooper’s hawks (
Accipiter cooperii
). Weasels (
Mustela
) including mink (
Mustela vison
) and also squirrels (
Sciuridae
) and black bears (
Ursus americanus
) have been reported to feed on eggs in nest boxes. Female buffleheads are particularly
vulnerable when perched on the nest, and eggs are vulnerable while females forage.
Ecosystem Roles
Buffleheads disperse seeds in their environment. They compete for nests with Barrow’s
goldeneyes (
Bucephala islandica
), common goldeneyes (
Bucephala clangula
), squirrels (
Sciuridae
), European starlings (
Sturnus vulgaris
) and northern flickers (
Colaptes auratus
).
Like many ducks, buffleheads are prone to infection by a variety of parasites. When
comparing buffleheads to other ducks, however, the abundance of parasitic species
is modest. By examining the gizzards and intestines of adult buffleheads several species
of
roundworms
,
flukes
,
tapeworms
, and
thorny-headed worms
were found. The
roundworm
species found were:
Amidostomum acutum
,
Capillaria anatis
,
Capillaria contorta
,
Echinuria parva
,
Ecinuria uncinata
,
Nematoda
Schistorophus
,
Streptocara crassicauda
,
Streptocara formosensis
,
Tetrameres crami
,
Tetrameres fissipina
, and
Tetrameres spinosa
. The
flukes
found were:
Apatemon canadensis
,
Apatemon gracilis
,
Cotylurus strigeoides
,
Dendritobilharzia pulverulenta
,
Echinoparypthium recurvatum
,
Echinostoma trivolvis
,
Gyrosoma marilae
,
Maritrema obstipum
,
Notochotylus attenuatus
,
Odhneria odhneri
,
Philophthalmus gralli
,
Plagiorchis elegans
,
Prosthogonimus cuneatus
,
Pseudosplotrema
,
Psilochasmus oxyurus
,
Strigea
, and
Zygocotyle lunata
. The
tapeworms
found were:
Abortilepis
,
Aploparaksis
,
Cloacotaenia
,
Cloacotaenia megalops
,
Dicranotaenia multisticta
,
Diorchis bulbodes
,
Diploposthe laevis
,
Fimbriaria
,
Gastrotaenia cygni
,
Hymenolepis
,
Lateriporus skrjabini
,
Microsomacanthus collaris
,
Microsomacanthus melanittae
,
Microsomacanthus parvula
,
Platyscolex ciliata
,
Retinometra albeola
, and
Shistocephalus
. Finally, the following
thorny-headed worms
found were:
Corynosoma constrictum
,
Polymorphus acutis
,
Polymorphus marilis
, and
Polymorphus obtusus
.
Of particular interest is that buffleheads appear to be the only duck with the tapeworm
Retinometra albeolae
. Leeches (
Theromyzon rude
,
Theromyzon tesulatum
, and
Theromyzon biannulatum
) may infest their upper respiratory tract as well as their eyes. Bufflehead ducklings
are more prone to leeches. A trematode in the family
Schistosomatidae
has been observed in the arteries. Renal coccidia also have been observed. Additionally,
avian cholera (
Pasteurella multocida
), avian malaria (
Plasmodium relictum
), and avian influenza all have been identified in buffleheads. The literature states
that little is known about the impacts of parasites and disease on buffleheads.
- Ecosystem Impact
- disperses seeds
- roundworms Amidostomum acutum
- roundworms Capillaria anatis
- roundworms Capillaria contorta
- roundworms Ecinuria parva
- roundworms Echinuria uncinata
- roundworms Schistorophus
- roundworms Streptocara crassicauda
- roundworms Streptocara formosensis
- roundworms Tetrameres crami
- roundworms Tetrameres fissipina
- roundworms Tetrameres spinosa
- flukes Apatemon canadensis
- flukes Apatemon gracillis
- flukes Cotylurus strigeoides
- flukes Dendritobilharzia pulverulenta
- flukes Echinoparypthium recurvatum
- flukes Echinostoma trivolvis
- flukes Gyrosoma marilae
- flukes Maritrema obstipum
- flukes Notochotylus attenuatus
- flukes Odhneria odhneri
- flukes Philophthalmus gralli
- flukes Plagiorchis elegans
- flukes Prosthogonimus cuneatus
- flukes Pseudosplotrema
- flukes Psilochasmus oxyurus
- flukes Strigea
- flukes Zygocotyle lunata
- tapeworms Abortilepis
- tapeworms Aploparaksis
- tapeworms Cloacotaenia
- tapeworms Cloacotaenia megalops
- tapeworms Dicranotaenia multisticta
- tapeworms Diorchis bulbodes
- tapeworms Diploposthe laevis
- tapeworms Fimbriaria
- tapeworms Gastrotaenia cygni
- tapeworms Hymenolepis
- tapeworms Lateriporus skrjabini
- tapeworms Microsomacanthus collaris
- tapeworms Microsomacanthus melanittae
- tapeworms Microsomacanthus parvula
- tapeworms Platyscolex ciliata
- tapeworms Retinometra albeola
- tapeworms Shistocephalus
- thorny-headed worms Corynosoma constrictum
- thorny-headed worms Polymorphus acutis
- thorny-headed worms Polymorphus marilis
- thorny-headed worms Polymorphus obtusus
Economic Importance for Humans: Positive
During the winter and fall months the habitat of buffleheads is prime duck hunting
range, so they are a target for hunters. Buffleheads make up 1 to 1.5% of all ducks
killed in the U.S. and 1.5 to 2% in Canada. During 2009, in the Atlantic Flyway of
Georgia, Maine and Maryland, 17,947 buffleheads were harvested. In 2008, there were
27,154 were harvested. Buffleheads also eat many types of insects, some of which are
pests to humans.
- Positive Impacts
- food
- controls pest population
Economic Importance for Humans: Negative
There are no adverse effects of Bucephala albeola on humans.
Conservation Status
Buffleheads are listed as least concern on the IUCN Red List and do not have special
status on US government lists. Previously, they were threatened by overshooting at
the end of the 19th and beginning 20th century. Toxic contaminants are a current threat
as well. Buffleheads collected around Long Island, NY, had low levels of polychlorinated
biphenyls (PCBs) and mercury in their bodies. However, when compared to other dangers,
habitat degradation is probably their biggest threat. Aspen nesting habitat has been
replaced over the last 100 years in western North America with agricultural land,
and clear-cutting for lumber continues to reduce the availability of nesting habitat.
Nest boxes have been installed in some areas to supplement nesting habitat. It is
important that these boxes be placed in conifer-heavy areas and the box openings be
the correct size and mimic their natural nesting preferences. These specifications
limit competition with other cavity-nesters.
Additional Links
Contributors
John Huth (author), Radford University, Karen Powers (editor), Radford University, Kiersten Newtoff (editor), Radford University, Melissa Whistleman (editor), Radford University, Catherine Kent (editor), Special Projects.
- Nearctic
-
living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- temperate
-
that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).
- terrestrial
-
Living on the ground.
- saltwater or marine
-
mainly lives in oceans, seas, or other bodies of salt water.
- freshwater
-
mainly lives in water that is not salty.
- taiga
-
Coniferous or boreal forest, located in a band across northern North America, Europe, and Asia. This terrestrial biome also occurs at high elevations. Long, cold winters and short, wet summers. Few species of trees are present; these are primarily conifers that grow in dense stands with little undergrowth. Some deciduous trees also may be present.
- tropical savanna and grassland
-
A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.
- savanna
-
A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.
- temperate grassland
-
A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.
- forest
-
forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.
- scrub forest
-
scrub forests develop in areas that experience dry seasons.
- coastal
-
the nearshore aquatic habitats near a coast, or shoreline.
- brackish water
-
areas with salty water, usually in coastal marshes and estuaries.
- marsh
-
marshes are wetland areas often dominated by grasses and reeds.
- swamp
-
a wetland area that may be permanently or intermittently covered in water, often dominated by woody vegetation.
- bog
-
a wetland area rich in accumulated plant material and with acidic soils surrounding a body of open water. Bogs have a flora dominated by sedges, heaths, and sphagnum.
- suburban
-
living in residential areas on the outskirts of large cities or towns.
- agricultural
-
living in landscapes dominated by human agriculture.
- riparian
-
Referring to something living or located adjacent to a waterbody (usually, but not always, a river or stream).
- estuarine
-
an area where a freshwater river meets the ocean and tidal influences result in fluctuations in salinity.
- monogamous
-
Having one mate at a time.
- iteroparous
-
offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).
- seasonal breeding
-
breeding is confined to a particular season
- sexual
-
reproduction that includes combining the genetic contribution of two individuals, a male and a female
- oviparous
-
reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.
- young precocial
-
young are relatively well-developed when born
- female parental care
-
parental care is carried out by females
- arboreal
-
Referring to an animal that lives in trees; tree-climbing.
- natatorial
-
specialized for swimming
- diurnal
-
- active during the day, 2. lasting for one day.
- nocturnal
-
active during the night
- crepuscular
-
active at dawn and dusk
- motile
-
having the capacity to move from one place to another.
- migratory
-
makes seasonal movements between breeding and wintering grounds
- territorial
-
defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement
- social
-
associates with others of its species; forms social groups.
- visual
-
uses sight to communicate
- acoustic
-
uses sound to communicate
- duets
-
to jointly display, usually with sounds in a highly coordinated fashion, at the same time as one other individual of the same species, often a mate
- food
-
A substance that provides both nutrients and energy to a living thing.
- carnivore
-
an animal that mainly eats meat
- insectivore
-
An animal that eats mainly insects or spiders.
- molluscivore
-
eats mollusks, members of Phylum Mollusca
- endothermic
-
animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.
- bilateral symmetry
-
having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.
- visual
-
uses sight to communicate
- tactile
-
uses touch to communicate
- acoustic
-
uses sound to communicate
- chemical
-
uses smells or other chemicals to communicate
References
Ahlund, M. 2005. Behavioural tactics at nest visits differ between parasites and hosts in brood-parasitic duck. Animal Behaviour , 70/2: 433-440.
Braune, B., B. Malone. 2006. Mercury and selenium in livers of waterfowl harvested in Northern Canada. Archives of Environmental Contamination and Toxicology , 50/2: 284-289.
Custer, C., T. Custer. 200. Organochlorine and trace element contamination in wintering and migrating diving ducks in the southern Great Lakes, USA, since the zebra mussel invasion. Environmental Toxicology and Chemistry , 19/11: 2821-2829.
Dugger, B., K. Dugger, L. Fredrickson. 2009. "The Birds of North America Online" (On-line). Accessed April 08, 2012 at http://bna.birds.cornell.edu/bna/species/098 .
Eadie, J., M. Mallory, H. Lumsden. 1995. "The Birds of North America Online" (On-line). Accessed April 08, 2012 at http://bna.birds.cornell.edu/bna/species/170 .
Evans, M., D. Lank, W. Boyd, F. Cooke. 2002. A comparison of the characteristics and fate of Barrow’s goldeneye and bufflehead nests in nest boxes and natural cavities. The Condor , 104/3: 610-619.
Ewart, M., J. McLaughlin. 1990. Helminths from spring and fall migrant bufflehead ducks ( Bucephala albeola ) at Delta, Manitoba, Canada. Canadian Journal of Zoology , 68/10: 2230-2233.
Gammonley, J., M. Heitmeyer. 1990. Behavior body condition, and foods of buffleheads and lesser scaups during spring migration through the Klamath Basin, California. Wilson Bulletin , 102/4: 672-683.
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