Campylopterus hemileucurusviolet sabrewing

Ge­o­graphic Range

Vi­o­let sabrew­ing hum­ming­birds (Campy­lopterus hemileu­cu­rus) have a broad ge­o­graphic range ex­tend­ing across the neotrop­ics. Vi­o­let sabrew­ings occur through­out por­tions of north­ern Mex­ico, El Sal­vador and Guatemala, to Costa Rica and Hon­duras. Their dis­tri­b­u­tion ex­tends as far south as north­ern South Amer­ica. ("IUCN Red List of Threat­ened Species", 2010; Sib­ley and Mon­roe Jr, 1990)

Habi­tat

Campy­lopterus hemileu­cu­rus is found in trop­i­cal habi­tats, par­tic­u­larly coastal slopes, in­land forests, and trop­i­cal grass­lands. They occur at el­e­va­tions rang­ing from 3,300 to 8,000 me­ters. Campy­lopterus hemileu­cu­rus does not mi­grate, be­cause food (nec­tar from flow­ers and small in­sects) is abun­dant in their habi­tats year-round. Thus, this re­gion pro­vides an ex­cel­lent place for breed­ing and there is no need for mi­gra­tion. (Land, 1963; Skutch, et al., 2010)

  • Range elevation
    3,300 to 8,000 m
    to ft

Phys­i­cal De­scrip­tion

Campy­lopterus hemileu­cu­rus is one of the larger hum­ming­bird species in the Fam­ily Trochil­i­dae, usu­ally grow­ing to an av­er­age length of 15 cm. It is bril­liantly col­ored, with a va­ri­ety of sharp vi­o­lets, greens, blacks, blues, and whites. The dark vi­o­let and bright blue feath­ers, mixed with dark for­est-green feath­ers, pro­vide ex­cel­lent cam­ou­flage in forested areas. Its name, sabrew­ing, refers to the strik­ing flat and thick shafts on its outer feath­ers. The bill is long and curved, well-adapted for ex­tract­ing nec­tar from flow­er­ing plants. Flow­ers with ra­dial sym­me­try are pre­ferred, be­cause they can eas­ily hover be­side the plant while ex­tract­ing nec­tar.

Male and fe­male Campy­lopterus hemileu­cu­rus have dif­fer­ent col­oration. The male's body is gen­er­ally dark vi­o­let and blue on the ven­tral side, with dark green or black on the dor­sal side. Fe­males tend to be more green­ish on the ven­tral side and black on the dor­sal side. Both are eas­ily rec­og­nized by their dis­tinc­tive vi­o­let throats. Males and fe­males share the same tail pat­tern with black and white col­oration. Ju­ve­nile vi­o­let sabrew­ings are dis­tin­guished by their lack of vi­o­let col­oration and flat feath­ers. (Land, 1963; Marin, 2001; Skutch, et al., 2010)

  • Sexual Dimorphism
  • sexes colored or patterned differently
  • male more colorful
  • Range mass
    9 to 12 g
    0.32 to 0.42 oz
  • Average length
    15.24 cm
    6.00 in
  • Average wingspan
    82.6 mm
    3.25 in

Re­pro­duc­tion

Campy­lopterus hemileu­cu­rus be­haves much like other species of hum­ming­birds dur­ing the mat­ing sea­son. Males form groups of six to ten and begin to sing loudly from their leks. Fe­males, upon hear­ing the mat­ing call, begin to build nests using mosses and other plant ma­te­r­ial. Grasses and small twigs found through­out the sur­round­ing habi­tat are used for nest build­ing just be­fore mat­ing. This usu­ally oc­curs dur­ing the rainy sea­son (May through Au­gust). Like all hum­ming­birds and typ­i­cally all lekking species, C. hemileu­cu­rus is polyg­y­nous. Pairs re­main to­gether only long enough for fer­til­iza­tion. The males then aban­don the nest, leav­ing the fe­males to in­cu­bate the eggs and care for the off­spring. (Marin, 2001)

The breed­ing sea­son for C. hemileu­cu­rus oc­curs dur­ing the rainy sea­son from May through Au­gust. It is thought that the hum­ming­birds choose this sea­son for its abun­dance of food, both for them­selves and for the off­spring. A clutch size of two eggs per nest is typ­i­cal. Fe­males in­cu­bate these eggs for 20 days. After a few hours of hatch­ing, fe­males begin feed­ing spi­ders and flu­ids to the off­spring. About 11 to 12 days later, young nestlings reach their full body mass, with males tend­ing to be larger than fe­males. Nestlings fledge 22 to 24 days after hatch­ing. In many hum­ming­bird species, the fe­male feeds her fledg­lings for 18 to 25 days after they have left the nest, but exact du­ra­tion for C. hemileu­cu­rus is un­known. Re­pro­duc­tive age is also cur­rently un­known.

Campy­lopterus hemileu­cu­rus usu­ally breeds twice per sea­son. Fe­males typ­i­cally build a sec­ond nest close to or on top of their first. (Marin, 2001; Schuch­mann, 1999)

  • Breeding interval
    Campylopterus hemileucurus breeds two times per season
  • Breeding season
    The breeding season occurs from May through August
  • Average eggs per season
    2
  • Average time to hatching
    21 days
  • Range fledging age
    22 to 24 days

Fe­male vi­o­let sabrew­ings are the main providers of care for off­spring. After the in­cu­ba­tion pe­riod, fe­males care for the young nestlings even after they have fledged. Within hours of being born, the moth­ers begin to feed the new­born nestlings a diet that con­sists of flu­ids and spi­ders. Dur­ing the com­ing weeks the mother will con­tinue to care for the young pro­tect­ing them from preda­tors such as other birds, mice, and cats until they have all reached in­de­pen­dence and are able to sur­vive on their own. (Marin, 2001; Schuch­mann, 1999)

  • Parental Investment
  • altricial
  • female parental care
  • pre-fertilization
    • provisioning
    • protecting
      • female
  • pre-hatching/birth
    • provisioning
      • female
    • protecting
      • female
  • pre-weaning/fledging
    • provisioning
      • female
    • protecting
      • female
  • pre-independence
    • provisioning
      • female
    • protecting
      • female

Lifes­pan/Longevity

Lit­tle or no data has been col­lected for lifes­pan of C. hemileu­cu­rus or other species in the genus Campy­lopterus. The av­er­age lifes­pan of hum­ming­birds in the Fam­ily Trochil­i­dae is 3 to 5 years. (Perry, 2011)

Be­hav­ior

Campy­lopterus hemileu­cu­rus is a soli­tary species that spends about three-fourths of its time perched. Like other hum­ming­birds, they are very cu­ri­ous and will in­ves­ti­gate all po­ten­tial food sources. They are di­ur­nal, with peak ac­tiv­ity dur­ing dawn and dusk. They ob­tain their food through­out the day. They hover in place while feed­ing by al­ter­nat­ing flap­ping their wings for­ward and back­ward. They can hover long enough to lap nec­tar from flow­ers. In flight, hum­ming­birds never stop beat­ing their wings. They can fly back­wards and up­side down, in ad­di­tion to their di­rect flight path. Dur­ing cold nights, hum­ming­birds go into tor­por in order to con­serve en­ergy. Through­out tor­por, their body tem­per­a­ture and heart­beat de­crease and they take on an ir­reg­u­lar breath­ing pat­tern.

Trop­i­cal hum­ming­birds, like vi­o­let sabrew­ings, do not mi­grate. How­ever, they change el­e­va­tion dur­ing sea­sonal and cli­matic changes.

Ter­ri­to­ri­al­ity is crit­i­cally im­por­tant among hum­ming­birds, es­pe­cially while for­ag­ing. Com­pared to smaller hum­ming­birds, like ruby-throated hum­ming­birds (Archilochus col­u­bris), vi­o­let sabrew­ings are not very ag­gres­sive. Due to their large size, they dom­i­nate dur­ing feed­ing and oth­ers often leave when they see this species ap­proach­ing. (Camp­bell and Lack, 1985; Gill, 1995; Sk­tuch and Singer, 1973; Tyrrell and Tyrell, 1985)

Home Range

Av­er­age ter­ri­tory size for this species is cur­rently un­known.

Com­mu­ni­ca­tion and Per­cep­tion

Campy­lopterus hemileu­cu­rus com­mu­ni­cates through calls and songs. Both males and fe­males pro­duce spe­cific calls. These calls are com­monly short sharp twit­ters made while vis­it­ing flow­ers to feed. Males gen­er­ate high-pitched songs that are used both to at­tract mates as well as to de­fend their ter­ri­to­ries. Like other hum­ming­birds (Fam­ily Trochil­i­dae), the wings of C. hemileu­cu­rus cre­ate a hum­ming noise dur­ing flight.

Campy­lopterus hemileu­cu­rus has ex­cel­lent vi­sual per­cep­tion and can see things at far dis­tances. They have color vi­sion and are drawn to brightly col­ored flow­ers for their nec­tar. They also have ul­tra-vi­o­let light per­cep­tion that aids in their for­ag­ing for nec­tar, since many flow­ers have such color pat­terns. They do not have a well-de­vel­oped sense of smell and gen­er­ally visit flow­ers with lit­tle to no scent. In ad­di­tion, their hear­ing is ex­tremely finely tuned. They can hear high-pitched sounds and de­tect tiny dif­fer­ences in sound qual­ity.

Like most birds, Campy­lopterus hemileu­cu­rus per­ceives its en­vi­ron­ment through vi­sual, tac­tile, au­di­tory and chem­i­cal stim­uli. (Camp­bell and Lack, 1985; How­ell, 2002)

Food Habits

Hum­ming­birds are very small birds with high me­tab­o­lisms. They must feed al­most con­stantly since most of their en­ergy is spent fly­ing. Campy­lopterus hemileu­cu­rus is pri­mar­ily nec­tiv­o­rous but also in­sec­tiv­o­rous. Most of their diet comes from flo­ral nec­tar, with the rest from arthro­pods (Phy­lum Arthro­poda), in­clud­ing flies (Order Diptera), spi­ders (Order Araneae), ants (Order Hy­menoptera), bee­tles (Order Coleoptera), and other small or­gan­isms. They oc­ca­sion­ally feed on non-in­sect arthro­pods. Their only lim­i­ta­tion in feed­ing is prey size, al­though they are ca­pa­ble of swal­low­ing sur­pris­ingly large or­gan­isms.

Food choice of all hum­ming­birds is chiefly de­ter­mined by sea­son and habi­tat. As a non-mi­gra­tory hum­ming­bird, C. hemileu­cu­rus de­pends on local food re­sources. They ob­tain nec­tar from brightly col­ored flow­ers, par­tic­u­larly those in the Neotrop­i­cal genus Mar­c­gravia dur­ing their flow­er­ing sea­son. They are most at­tracted to red and yel­low flow­ers that are shaped like their beaks (long, tubu­lar, and ra­di­ally sym­met­ric). When hum­ming­birds open their beaks, they lap up the nec­tar with their tongues, which have grooves on the sides that col­lect the liq­uid. Vi­o­let sabrew­ings can con­sume con­sid­er­able amounts of nec­tar, al­most equal to twice their weight, on a daily basis.

Con­ve­nience also plays a major role in the feed­ing pat­terns of this bird. They visit flow­ers from which they can most eas­ily ob­tain in­sects and nec­tar. Dur­ing other parts of the year, when flo­ral nec­tar is lim­ited, arthro­pods are their main food source. Dur­ing this pe­riod, C. hemileu­cu­rus often hov­ers over for­est streams and darts at large swarms of gnats. This hov­er­ing tech­nique dur­ing feed­ing is com­mon in all hum­ming­birds. (Rem­sen, et al., 1986; Wag­ner, 1946)

  • Animal Foods
  • insects
  • terrestrial non-insect arthropods
  • Plant Foods
  • nectar

Pre­da­tion

Adult vi­o­let sabrew­ings have few preda­tors. This has been at­trib­uted to mostly to their large body size. How­ever, ju­ve­nile hum­ming­birds are threat­ened by mice and cats. Nest preda­tors are the great­est threat to off­spring; these in­clude snakes, jays, tou­cans, hawks, and a few bats.

Male vi­o­let sabrew­ings are easy to rec­og­nize with their bright pur­ple col­or­ing and large size. Fe­males, in con­trast, have a cryp­tic col­oration, an adap­ta­tion that cam­ou­flages them from preda­tors. Fe­male hum­ming­birds also fly in a zigzag when re­turn­ing to their nests in order to evade preda­tors. (Camp­bell and Lack, 1985; Sk­tuch and Singer, 1973)

  • Anti-predator Adaptations
  • cryptic

Ecosys­tem Roles

Campy­lopterus hemileu­cu­rus, like many other species of hum­ming­birds, are pol­li­na­tors. They pol­li­nate var­i­ous trop­i­cal plants while feed­ing on nec­tar. Of­ten­times they pol­li­nate plants used to shade cof­fee plan­ta­tions. With­out these birds and their rel­a­tives, many trop­i­cal plants would be un­able to re­pro­duce and local cof­fee pro­duc­tion would be re­duced. A small group of in­ver­te­brates known as hum­ming­bird mites also feed on nec­tar. They use hum­ming­birds to trans­port them from plant to plant to feed on nec­tar. (Cof­fee & Con­ser­va­tion, 2009)

  • Ecosystem Impact
  • pollinates
Com­men­sal/Par­a­sitic Species
  • Hum­ming­bird mites

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Campy­lopterus hemileu­cu­rus plays a crit­i­cal role in pol­li­na­tion of neotrop­i­cal plants. It pol­li­nates a va­ri­ety of trop­i­cal plants, but the most im­por­tant to hu­mans are trees in the genus (Car­po­da­cus). These trees are im­por­tant in pro­vid­ing shade for cof­fee plan­ta­tions. (Cof­fee & Con­ser­va­tion, 2009)

  • Positive Impacts
  • pollinates crops

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

There are no known ad­verse af­fects of C. hemileu­cu­rus on hu­mans.

Con­ser­va­tion Sta­tus

Based on the IUCN Red List, the con­ser­va­tion sta­tus of C. hemileu­cu­rus is "Least Con­cern". Pop­u­la­tions are not be­lieved to be de­creas­ing rapidly enough to ap­proach the thresh­olds for "Vul­ner­a­ble" sta­tus.

The major threats to hum­ming­birds are habi­tat degra­da­tion, loss, and frag­men­ta­tion. Al­though vi­o­let sabrew­ings are not listed, most North Amer­i­can species are pro­tected under the Mi­gra­tory Bird Treaty Act. All hum­ming­bird species are listed by CITES in Ap­pen­dix II ex­cept the hook-billed hum­ming­bird (Glau­cis dohrnii), which is listed in Ap­pen­dix I. ("IUCN Red List of Threat­ened Species", 2010; CITES, 2010; Michi­gan Nat­ural Fea­tures In­ven­tory, 2009; U.S. Fish & Wildlife Ser­vice, 2011)

Con­trib­u­tors

Patrick Boyd (au­thor), Rad­ford Uni­ver­sity, Heather Vin­ing (au­thor), Rad­ford Uni­ver­sity, Chris­tine Small (ed­i­tor), Rad­ford Uni­ver­sity, Rachelle Ster­ling (ed­i­tor), Spe­cial Pro­jects.

Glossary

Neotropical

living in the southern part of the New World. In other words, Central and South America.

World Map

acoustic

uses sound to communicate

altricial

young are born in a relatively underdeveloped state; they are unable to feed or care for themselves or locomote independently for a period of time after birth/hatching. In birds, naked and helpless after hatching.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

chemical

uses smells or other chemicals to communicate

coastal

the nearshore aquatic habitats near a coast, or shoreline.

cryptic

having markings, coloration, shapes, or other features that cause an animal to be camouflaged in its natural environment; being difficult to see or otherwise detect.

diurnal
  1. active during the day, 2. lasting for one day.
endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

female parental care

parental care is carried out by females

forest

forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.

herbivore

An animal that eats mainly plants or parts of plants.

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

motile

having the capacity to move from one place to another.

native range

the area in which the animal is naturally found, the region in which it is endemic.

nectarivore

an animal that mainly eats nectar from flowers

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

polygynous

having more than one female as a mate at one time

seasonal breeding

breeding is confined to a particular season

sedentary

remains in the same area

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

solitary

lives alone

tactile

uses touch to communicate

terrestrial

Living on the ground.

territorial

defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement

tropical

the region of the earth that surrounds the equator, from 23.5 degrees north to 23.5 degrees south.

tropical savanna and grassland

A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.

savanna

A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.

temperate grassland

A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.

visual

uses sight to communicate

Ref­er­ences

2010. "IUCN Red List of Threat­ened Species" (On-line). Campy­lopterus hemileu­cu­rus. Ac­cessed April 04, 2011 at www.​iucnredlist.​org.

Bames­berger, J., V. Fitz, J. At­wood. 1991. Tem­po­ral pat­terns of singing ac­tiv­ity at leks of the White-Bel­lied Emer­ald. The Wil­son Bul­letin, 103/3: 373-386.

Bird, D. 2004. The Bird Al­manac: A Guide to Es­sen­tial Facts and Fig­ures of the World's Birds. New York: Fire­fly Books.

CITES, 2010. "Con­ven­tion on In­ter­na­tional Trade in En­dan­gered Species of Wild Fauna and Flora" (On-line). Ac­cessed April 04, 2011 at http://​www.​cites.​org/​eng/​app/​appendices.​shtml.

Camp­bell, B., E. Lack. 1985. A Dic­tio­nary of Birds. Vemil­lion: Buteo Books.

Cof­fee & Con­ser­va­tion, 2009. "Cof­fee and con­ser­va­tion are your beans for the birds" (On-line). Ac­cessed April 05, 2011 at http://​www.​coffeehabitat.​com/​2009/​09/​violet-sabrewing/​.

Dickey, D., A. Van Rossem. 1938. The Birds of El Sal­vador. Chicago: Field Mu­seum of Na­tional His­tory.

Dun­ning, J. 2008. CRC Hand­book of Avian Body Masses. Florida: CRC Press.

Gill, F. 1995. Or­nithol­ogy, Sec­ond Edi­tion. New York: Co­lum­bia Uni­ver­sity Press.

Hen­der­son, C. 2010. Birds of Costa Rica: A Field Guide. Texas: Uni­ver­sity of Texas Press.

How­ell, S. 2002. Hum­ming­birds of North Amer­ica. San Diego: AP Nat­ural World.

Land, H. 1963. A col­lec­tion of birds from the Caribbean low­lands of Guatemala. The Con­dor, 65/1: 49-65.

Marin, M. 2001. Post­na­tal de­vel­op­ment of the vi­o­let sabrew­ing in Costa Rica. The Wil­son Bul­letin, 113/1: 110-114.

Michi­gan Nat­ural Fea­tures In­ven­tory, 2009. "Michi­gan's Spe­cial An­i­mals" (On-line). Ac­cessed April 04, 2011 at http://​web4.​msue.​msu.​edu/​mnfi/​data/​specialanimals.​cfm#​grp3.

Perry, L. 2011. "Those hum­ming­birds in your gar­den" (On-line). Ac­cessed May 01, 2011 at http://​www.​uvm.​edu/​pss/​ppp/​articles/​hummers2.​html.

Rem­sen, J., F. Stiles, P. Scott. 1986. Fre­quency of arthro­pods in stom­achs of trop­i­cal hum­ming­birds. The Auk, 103/2: 436-441.

Ridg­way, R., H. Fried­mann. 1911. The Birds of North and Mid­dle Amer­ica. Wash­ing­ton: Gov­ern­ment print­ing of­fice.

Schuch­mann, K. 1999. Fam­ily Trochil­i­dae. J del Hoyo, A El­liott, S Jordi, eds. Hand­book of the Birds of the World, Vol. 5. Barcelona: Lynx Edi­cions.

Sib­ley, C., B. Mon­roe Jr. 1990. Dis­tri­b­u­tion and Tax­on­omy of Birds of the World. New Haven: Yale Uni­ver­sity Press.

Sk­tuch, A., A. Singer. 1973. The Life of the Hum­ming­bird. New York: Crown Pub­lish­ers, Inc..

Skutch, A., S. Adams, C. Hen­der­son. 2010. Birds of Costa Rica: A Field Guide. Austin Texas: Uni­ver­sity of Texas Press.

Tyrrell, E., R. Tyrell. 1985. Hum­ming­birds: Their Life and Be­hav­ior. New York: Crown Pub­lish­ers, Inc.

U.S. Fish & Wildlife Ser­vice, 2011. "Mi­gra­tory Bird Pro­gram" (On-line). Ac­cessed April 04, 2011 at http://​www.​fws.​gov/​migratorybirds/​.

Wag­ner, H. 1946. Food and feed­ing habits of Mex­i­can hum­ming­birds. The Wil­son Bul­letin, 58/2: 69-93.

Williamson, S. 2001. A Field Guide to Hum­ming­birds of North Amer­ica. New York: Houghton Mif­flin Com­pany.

Willmer, P., S. Cor­bet. 1981. Tem­po­ral and mi­cro­cli­matic par­ti­tion­ing of the flo­ral re­sources of Jus­ti­cia aurea amongst a con­course of pollen vec­tors and nec­tar rob­bers. Oe­colo­gia, 51/1: 67-78.