Nestor notabiliskea

Ge­o­graphic Range

Kea are found only in the moun­tains of South Is­land, New Zealand. (del Hoyo, et al., 1996; Teb­bich, et al., 1996)

Habi­tat

Kea live in wooded val­leys and south­ern beech (Nothofa­gus) forests that line sub-alpine scrub­lands at 600 to 2000 m. In sum­mer, kea in­habit high el­e­va­tion scrub and alpine tun­dra areas. In au­tumn, they move to higher el­e­va­tions to for­age for berries. In win­ter, kea move below the tim­ber­line. (del Hoyo, et al., 1996)

  • Range elevation
    600 to 2000 m
    1968.50 to 6561.68 ft

Phys­i­cal De­scrip­tion

Kea are crow-sized par­rots, about 48 cm long as adults. They have brown­ish-green heads and un­der­parts with black­ish edges. Their bod­ies have dull bronze-green plumage. The outer webs of their pri­maries are dull blue, and the un­der­wing coverts are or­ange red with yel­low bar­ring and notch­ing that ex­tends to the un­der­sides of the flight feath­ers. The lower back is dull red in color, reach­ing to the up­per­tail coverts. The upper sur­face of the tail is bronze-green, and the under sur­face of the tail is dull yel­low. Kea have de­curved upper bills (cul­mens). Fe­males have shorter, less curved cul­mens and weigh about 20 per­cent less than males. Ju­ve­nile kea have yel­low­ish crowns and ceres. (Bond, et al., 1991; del Hoyo, et al., 1996; Di­a­mond and Bond, 1999)

  • Sexual Dimorphism
  • male larger
  • Average mass
    922 g
    32.49 oz
  • Average length
    48 cm
    18.90 in

Re­pro­duc­tion

Kea have a polyg­y­nous mat­ing sys­tem. Males fight for dom­i­nance, and the hi­er­ar­chy is strict: as few as 10% of males may be al­lowed to breed in cer­tain years. Cop­u­la­tion is often ini­ti­ated the fe­male, who ap­proaches the male and in­vites play or adopts a sub­mis­sive pos­ture and so­lic­its preen­ing. The male then feeds the fe­male a re­gur­gi­tated meal and mounts her. (del Hoyo, et al., 1996; Di­a­mond and Bond, 1999; Teb­bich, et al., 1996)

Kea have been ob­served breed­ing at all times of the year, ex­cept late au­tumn. Their main re­pro­duc­tive pe­riod lasts from July to Jan­u­ary. They nest in bur­rows under rocks or among tree roots. Kea have clutches of two to four eggs, and in­cu­bate the eggs for three to four weeks. The al­tri­cial hatch­lings fledge after 13 weeks, and then dis­perse from their natal ranges after an­other five to six weeks. Males are sex­u­ally ma­ture after four or five years, while fe­males be­come sex­u­ally ma­ture as early as three years of age. (del Hoyo, et al., 1996; Di­a­mond and Bond, 1999; Teb­bich, et al., 1996)

  • Breeding interval
    Kea breed once yearly.
  • Breeding season
    Kea breed mainly between January and July.
  • Range eggs per season
    2 to 4
  • Range time to hatching
    3 to 4 weeks
  • Average fledging age
    13 weeks
  • Range time to independence
    18 to 19 weeks
  • Range age at sexual or reproductive maturity (female)
    3 (low) years
  • Range age at sexual or reproductive maturity (male)
    4 to 5 years

One a fe­male kea lays her eggs, she sits on the nest and in­cu­bates them for three weeks. Dur­ing this time, she rarely leaves the nest and the male feeds her. After the eggs hatch, the male con­tin­ues to feed the fe­male, and she, in turn, feeds the chicks. After a month, the male be­gins feed­ing the chicks him­self. The chicks fledge at 9 to 13 weeks of age, and the male as­sumes sole re­spon­si­bil­ity for feed­ing them. He con­tin­ues feed­ing his fledg­lings for up to six weeks. Af­ter­ward, the ju­ve­niles dis­perse from their natal area and travel to­gether in flocks for two to three years be­fore set­tling down. (Di­a­mond and Bond, 1999)

  • Parental Investment
  • altricial
  • pre-fertilization
    • protecting
      • female
  • pre-hatching/birth
    • provisioning
      • female
    • protecting
      • female
  • pre-weaning/fledging
    • provisioning
      • male
      • female
    • protecting
      • male
      • female
  • pre-independence
    • provisioning
      • male
    • protecting
      • male
      • female
  • extended period of juvenile learning

Lifes­pan/Longevity

Kea can live 14.4 years in cap­tiv­ity. Life span in the wild has not been re­ported. (Carey and Judge, 2002)

Be­hav­ior

Kea are highly in­tel­li­gent, so­cial birds. They live in fam­ily groups, and ag­gre­ga­tions of 30 to 40 birds often for­age at prime feed­ing grounds, such as garbage dumps. Kea ex­hibit a va­ri­ety of so­cial be­hav­iors, in­clud­ing in­tri­cate play. They have dom­i­nance hi­er­ar­chies, but these hi­er­ar­chies are not nec­es­sar­ily lin­ear. For ex­am­ple, an adult male may be dom­i­nant to a subadult male, who is dom­i­nant to a ju­ve­nile male, who, in turn, is dom­i­nant to the adult male. Ex­per­i­ments on co­op­er­a­tion in kea sug­gest that dom­i­nant in­di­vid­u­als can force sub­or­di­nants to co­op­er­ate in tasks that ben­e­fit only the dom­i­nant birds. Also, it has been shown that kea in cap­tiv­ity can learn com­pli­cated tasks from ob­serv­ing oth­ers, though this abil­ity has not been shown for kea in the wild. It has been pro­posed that life in an ex­treme alpine en­vi­ron­ment has en­cour­aged kea to op­por­tunis­ti­cally ex­plore their sur­round­ings. They com­monly in­ves­ti­gate human be­long­ings, and are known to de­stroy car ac­ces­sories and ski lodge equip­ment. Kea are di­ur­nal, ris­ing in the early morn­ing to begin call­ing and then for­ag­ing until late morn­ing. They gen­er­ally roost dur­ing the mid­dle of the day, and begin for­ag­ing again in the evening, some­times until after dark, when they go to roost for the night on tree branches. The tim­ing of these daily ac­tiv­i­ties varies with the weather; kea are fairly heat-in­tol­er­ant and spend more time roost­ing on hot days. (Di­a­mond and Bond, 1999; Gaj­don, et al., 2004; Huber and Taborsky, 2001; Teb­bich, et al., 1996)

Home Range

Home range size in kea are not re­ported.

Com­mu­ni­ca­tion and Per­cep­tion

Kea per­ceive vi­sual, tac­tile, au­di­tory, and chem­i­cal stim­uli. They com­mu­ni­cate with a wide repetoire of vo­cal­iza­tions, in­clud­ing the "kee-ah" flight call for which they are named. They also com­mu­ni­cate by fluff­ing their head feath­ers into var­i­ous "fa­cial ex­pres­sions" and by pos­tur­ing. (Di­a­mond and Bond, 1999)

Food Habits

Kea are op­por­tunis­tic, om­niv­o­rous par­rots. The leaves, buds, and nuts of south­ern beeches (Nothofa­gus) are es­pe­cially im­por­tant in the kea diet. The foods con­sumed vary by sea­son, how­ever. In spring they eat moun­tain daisies (Celmisia) and dig in the soil for small plants and in­sects. In sum­mer kea con­sume the nec­tar and pollen of flow­er­ing moun­tain flax (Phorium colen­soi) and rata (Met­rosideros). They eat berries of co­prosma (Co­prosma) and snow to­tara (Podocar­pus ni­valis), and eat the leaves, fruit, seeds, and flow­ers of other plants. In sum­mer they also eat bee­tle grubs, grasshop­pers, and land snails. In fall kea feed on moun­tain beech leaves and buds and con­tinue for­ag­ing on the roots, bulbs, fruit, seeds, and stems of other plants. Kea scav­enge on trash heaps year round and rel­ish the flesh and bone mar­row from car­casses. These food sources be­come par­tic­u­larly im­por­tant in win­ter, when plant foods are scarce. Fi­nally, kea have been re­ported to eat rab­bits and mice, and they have gained a rep­u­ta­tion for at­tack­ing sheep, al­though they usu­ally only prey on wounded or dis­eased sheep. (del Hoyo, et al., 1996; Di­a­mond and Bond, 1999; Math­ew­son, 1991)

  • Animal Foods
  • mammals
  • carrion
  • insects
  • mollusks
  • Plant Foods
  • leaves
  • roots and tubers
  • seeds, grains, and nuts
  • fruit
  • nectar
  • pollen
  • flowers

Pre­da­tion

New Zealand fal­cons (Falco no­vaezee­landiae) have been ob­served at­tack­ing kea, but no one has re­ported an in­ci­dence of suc­cess­ful pre­da­tion. Kea re­main alert for air at­tacks when for­ag­ing, and they band to­gether to chase fal­cons that threaten a mem­ber of their group. (Di­a­mond and Bond, 1999)

Ecosys­tem Roles

Kea, being op­por­tunis­tic, gen­er­al­ist for­agers, are pri­mary, sec­ondary, and higher-level con­sumers. In the past, kea prob­a­bly had an array of com­peti­tors, such as kaka (Nestor merid­ion­alis), moa (Anom­alopteryx, Di­nor­nis, Emeus, Eu­ryapteryx, Mega­lapteryx, and Pachy­or­nis spp.), kakapo (Strigops habrop­tila), takahe (Por­phyrio man­telli), and New Zealand ravens (Corvus mori­o­rum). But human set­tle­ment fu­eled a mass ex­tinc­tion of New Zealand's na­tive birds. Moa, takahe, and New Zealand ravens are now ex­tinct, and kakapo are ex­tremely rare. Only kaka re­main to com­pete with kea and, where their ranges over­lap, these two closely re­lated species use many of the same food re­sources. (Di­a­mond and Bond, 1999)

Mu­tu­al­ist Species

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Kea are im­por­tant for New Zealand's tourism in­dus­try. These birds have been called "the clown of New Zealand's South­ern Alps" by the De­part­ment of Con­ser­va­tion, at­tract­ing crowds when they con­vene on au­to­mo­biles. (Di­a­mond and Bond, 1999; Math­ew­son, 1991)

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

Kea have been known to at­tack sheep, and the wounds can be­come in­fected with Clostrid­ium bac­te­ria. The bac­te­ria can cause blood poi­son­ing, which can be fatal to sheep. In­creas­ingly, the par­rots have come into con­tact with human habi­ta­tions, some­times for­ag­ing at dumps and cab­ins. Kea have been known to de­stroy car ac­ces­sories, such as wind­shield wipers and weather strip­ping. These birds also have shred­ded hik­ing boots and have stolen ob­jects such as sun­glasses. The dam­age can cause se­ri­ous prob­lems, such as when the birds rip out car wiring and de­stroy ski-lift warn­ing sys­tems. (Di­a­mond and Bond, 1999)

Con­ser­va­tion Sta­tus

Kea are cur­rently clas­si­fied as vul­ner­a­ble by the IUCN, and they are a BirdLife "re­stricted-range" species. They are sub­ject to in­ter­na­tional trade reg­u­la­tions under CITES ap­pen­dix II, as are most par­rots. Kea are also pro­tected within New Zealand by the Wildlife Act of 1953, the Na­tional Parks Act, the An­i­mals Pro­tec­tion Act, and the Trade in En­dan­gered Species Act. These laws pro­hibit the cap­ture of kea on pri­vate and pub­lic lands, pro­hibit their mis­treat­ment, and ban their ex­port. How­ever, par­rot-smug­gling is a lu­cra­tive busi­ness, and kea are often cap­tured and ex­ported for the black mar­ket pet trade. It is un­known ex­actly how many kea are left in the wild. Es­ti­mates range from only 2,000 to 5,000 birds, but for now, kea pop­u­la­tions ap­pear to be sta­ble--es­pe­cially in na­tional parks and other pro­tected areas. (del Hoyo, et al., 1996; Di­a­mond and Bond, 1999)

Con­trib­u­tors

Al­li­son Poor (), Uni­ver­sity of Michi­gan-Ann Arbor.

Al­li­son Poor (au­thor), Uni­ver­sity of Michi­gan-Ann Arbor.

Re­becca Williams (au­thor), Uni­ver­sity of Michi­gan-Ann Arbor, Terry Root (ed­i­tor), Uni­ver­sity of Michi­gan-Ann Arbor.

Glossary

Australian

Living in Australia, New Zealand, Tasmania, New Guinea and associated islands.

World Map

acoustic

uses sound to communicate

altricial

young are born in a relatively underdeveloped state; they are unable to feed or care for themselves or locomote independently for a period of time after birth/hatching. In birds, naked and helpless after hatching.

arboreal

Referring to an animal that lives in trees; tree-climbing.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

carrion

flesh of dead animals.

causes or carries domestic animal disease

either directly causes, or indirectly transmits, a disease to a domestic animal

chemical

uses smells or other chemicals to communicate

diurnal
  1. active during the day, 2. lasting for one day.
dominance hierarchies

ranking system or pecking order among members of a long-term social group, where dominance status affects access to resources or mates

ecotourism

humans benefit economically by promoting tourism that focuses on the appreciation of natural areas or animals. Ecotourism implies that there are existing programs that profit from the appreciation of natural areas or animals.

endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

forest

forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.

island endemic

animals that live only on an island or set of islands.

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

migratory

makes seasonal movements between breeding and wintering grounds

motile

having the capacity to move from one place to another.

mountains

This terrestrial biome includes summits of high mountains, either without vegetation or covered by low, tundra-like vegetation.

native range

the area in which the animal is naturally found, the region in which it is endemic.

omnivore

an animal that mainly eats all kinds of things, including plants and animals

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

polygynous

having more than one female as a mate at one time

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

social

associates with others of its species; forms social groups.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

terrestrial

Living on the ground.

visual

uses sight to communicate

year-round breeding

breeding takes place throughout the year

Ref­er­ences

Bond, A., K. Wil­son, J. Di­a­mond. 1991. Sex­ual di­mor­phism in the kea (Nestor no­ta­bilis). Emu, 91: 12-19.

Carey, J., D. Judge. 2002. "Longevity records: Life spans of mam­mals, birds, am­phib­ians, rep­tiles, and fish" (On-line). Max Planck In­sti­tute for De­mo­graphic Re­search. Ac­cessed Oc­to­ber 03, 2005 at http://​www.​demogr.​mpg.​de/​.

Di­a­mond, J., A. Bond. 1999. Kea, bird of para­dox: the evo­lu­tion and be­hav­ior of a New Zealand par­rot. Berke­ley, CA: Uni­ver­sity of Cal­i­for­nia Press, Ltd..

Gaj­don, G., N. Fijn, L. Huber. 2004. Test­ing so­cial learn­ing in a wild moun­tain par­rot, the kea (Nestor no­ta­bilis). Learn­ing & Be­hav­ior, 32 (1): 62-71.

Huber, L., M. Taborsky. 2001. So­cial ef­fects on ob­ject-ex­plo­ration in keas. Ad­vances in Ethol­ogy, 36: 181-182.

Math­ew­son, W. 1991. Cop­ing with the killer kea. In­ter­na­tional Wildlife, 21: 36-7.

Teb­bich, S., M. Taborsky, H. Win­kler. 1996. So­cial ma­nip­u­la­tion causes co­op­er­a­tion in keas. An­i­mal Be­hav­iour, 52: 1-10.

del Hoyo, J., A. El­liot, J. Sar­gatal. 1996. Hand­book of the Birds of the World: Vol­ume 4: Sand­grouse to Cuck­oos. Barcelona: Lynx Edi­cions.