Rodolia cardinalisvedalia

Ge­o­graphic Range

Al­though na­tive to Aus­tralia, Rodolia car­di­nalis flour­ishes today through­out Aus­tralia, the United States, and Eu­rope in areas where cit­rus is grown. (Bor­ror, et al., 1981; Wee­den, et al., 2004)

Habi­tat

Rodolia car­di­nalis are spe­cial­ists, feed­ing on Icerya pur­chasi which in turn spe­cial­ize on cit­rus trees. For this rea­son, Rodolia car­di­nalis live in cit­rus or­chards and nearby hills and scrub­land in areas where Icerya pur­chasi are also found. In ad­di­tion to cit­rus, cot­tony cus­sion scales are found on Aca­cia, box­wood, cit­rus, mag­no­lia, Nan­d­ina, olive, Pit­tospo­rum, and rose plants, which there­for ad­di­tion­ally pro­vide habi­tat for the vedalia bee­tle. (Buchs­baum, et al., 1987; Wee­den, et al., 2004)

Phys­i­cal De­scrip­tion

This species of la­dy­bee­tle, like oth­ers, has a very dis­tinc­tive shape. The body is broadly oval to nearly spher­i­cal, and is strongly con­vex dor­sally and nearly flat ven­trally. The head has short an­ten­nae and is partly or com­pletely con­cealed by the prono­tum. Fe­males are very red in color, while males are more black al­though fine body hairs often ob­scure the color pat­tern and give a de­cid­edly grey ap­pear­ance. Eggs are bright red. Ma­ture lar­vae are pink­ish with black mark­ings and often have a blueish tinge. (Bor­ror and White, 1970; Essig, 1926)

  • Sexual Dimorphism
  • sexes colored or patterned differently
  • female more colorful

Re­pro­duc­tion

Rodolia car­di­nalis re­pro­duces sex­u­ally. In warm cli­mates there are sev­eral gen­er­a­tions each year. The bright red eggs are laid on the egg sac of the cot­tony cush­ion scale in small, closely packed masses of one or two dozen. Upon hatch­ing, the small pink­ish lar­vae im­me­di­ately enter the sac and feed on the scale eggs. These lar­vae are usu­ally found in aphid colonies. (Bor­ror and White, 1970; Milne and Milne, 1980; Wald­bauer, 2000)

  • Range eggs per season
    150 to 190

Be­hav­ior

Vedalia bee­tles are com­monly seen on plants and often over­win­ter as adults in large swarms under fallen leaves or bark. Win­ter­ing ag­gre­ga­tions are col­lected by the bushel from high Cal­i­for­nia hills and sold to cit­rus grow­ers. In early spring the bee­tles are lib­er­ated in or­chards; oth­ers make their own way down into val­leys where cit­rus is grown. Dur­ing the dry sea­son, new gen­er­a­tions re­turn to a high, cool refuge. The suc­cess of the in­sect is due to the ra­pid­ity of de­vel­op­ment, har­di­ness, and their greedy ap­petites. (Bor­ror, et al., 1981; Buchs­baum, et al., 1987; Milne and Milne, 1980)

Food Habits

Both adults and lar­vae are vo­ra­cious preda­tors. Ho­mopteran pests in­clud­ing aphids, scale in­sects, mealy bugs, and mites are food for R. car­di­nalis. Adults and lar­vae search the fo­liage of bushes and trees in search of prey. (Buchs­baum, et al., 1987; Bur­ton, 1968)

  • Animal Foods
  • insects

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Rodolia car­di­nalis is used com­mer­cially to com­bat scale in­sects in­ju­ri­ous to cit­rus or­chards, specif­i­cally Icerya pur­chasi, the cot­tony cush­ion scale. Or­chard own­ers use this form of In­te­grated Pest Man­age­ment(IPM)--use of a va­ri­ety of strate­gies for the con­trol of in­sect pests, in­clud­ing cul­tural, chem­i­cal, and bi­o­log­i­cal con­trols while re­strict­ing and al­ter­ing the use of pes­ti­cides--to con­trol a pest pop­u­la­tion al­ways pre­sent. The nat­ural preda­tor of Icerya Pur­chasi is used (R.​cardin­alis) in­stead of pes­ti­cides and other such ma­te­ri­als. Ag­gre­ga­tions of the bee­tle are col­lected by the bushel and sold to cit­rus grow­ers. When im­ported from Aus­tralia to Cal­i­for­nia in the 19th cen­tury,they brought with them no nat­ural preda­tors, mak­ing them an ideal an­i­mal to use for pest man­age­ment. (Bel­lamy and Evans, 1996; Bur­ton, 1968)

  • Positive Impacts
  • controls pest population

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

In its North Amer­i­can habi­tat, the vedalia bee­tle has no nat­ural preda­tors and so can out­com­pete na­tive la­dy­bugs which also offer im­por­tant pest con­trol ben­e­fits.

Con­ser­va­tion Sta­tus

Rodolia car­di­nalis is ex­tremely sen­si­tive to some in­sec­ti­cides such as Baythroid, com­monly used on cit­rus crops. Global pop­u­la­tions are strong how­ever, and re­quire no spe­cial con­ser­va­tion sta­tus. (Wee­den, et al., 2004)

Other Com­ments

In the Mid­dle Ages, cer­tain species of coc­cinel­lid bee­tles were ded­i­cated to the Vir­gin Mary and named "bee­tles of Our Lady." As time pro­gressed la­dy­bird bee­tles, la­dy­birds, or la­dy­bugs be­came pop­u­lar names with Eng­lish-speak­ing chil­dren.

Rodolia car­di­nalis is na­tive to Aus­tralia. How­ever, in 1868 Icerya pur­chasi, a cot­tony cush­ion scale (and also from Aus­tralia), was in­tro­duced to Cal­i­for­nia ac­ci­den­tally as a se­ri­ous pest of cit­rus in Cal­i­for­nia. In 1888-1889 R. car­di­nalis was in­ten­tion­ally in­tro­duced to Cal­i­for­nia be­cause it was a nat­ural preda­tor of the scale. In less than 2 years the scale was under com­plete con­trol and R. car­di­nalis adapted and flour­ished in the new en­vi­ron­ment. Its dis­tri­b­u­tion now in­cludes Aus­tralia, the United States, and Eu­rope. (Bel­lamy and Evans, 1996; Bor­ror, et al., 1981; Wee­den, et al., 2004)

Con­trib­u­tors

Sara Di­a­mond (ed­i­tor), An­i­mal Di­ver­sity Web.

Heather Mar­tin (au­thor), South­west­ern Uni­ver­sity, Stephanie Fab­ri­tius (ed­i­tor), South­west­ern Uni­ver­sity.

Glossary

Australian

Living in Australia, New Zealand, Tasmania, New Guinea and associated islands.

World Map

Nearctic

living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.

World Map

Palearctic

living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.

World Map

agricultural

living in landscapes dominated by human agriculture.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

carnivore

an animal that mainly eats meat

colonial

used loosely to describe any group of organisms living together or in close proximity to each other - for example nesting shorebirds that live in large colonies. More specifically refers to a group of organisms in which members act as specialized subunits (a continuous, modular society) - as in clonal organisms.

diurnal
  1. active during the day, 2. lasting for one day.
ectothermic

animals which must use heat acquired from the environment and behavioral adaptations to regulate body temperature

fertilization

union of egg and spermatozoan

heterothermic

having a body temperature that fluctuates with that of the immediate environment; having no mechanism or a poorly developed mechanism for regulating internal body temperature.

hibernation

the state that some animals enter during winter in which normal physiological processes are significantly reduced, thus lowering the animal's energy requirements. The act or condition of passing winter in a torpid or resting state, typically involving the abandonment of homoiothermy in mammals.

insectivore

An animal that eats mainly insects or spiders.

internal fertilization

fertilization takes place within the female's body

introduced

referring to animal species that have been transported to and established populations in regions outside of their natural range, usually through human action.

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

native range

the area in which the animal is naturally found, the region in which it is endemic.

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

suburban

living in residential areas on the outskirts of large cities or towns.

terrestrial

Living on the ground.

tropical

the region of the earth that surrounds the equator, from 23.5 degrees north to 23.5 degrees south.

tropical savanna and grassland

A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.

savanna

A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.

temperate grassland

A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.

year-round breeding

breeding takes place throughout the year

Ref­er­ences

Bel­lamy, C., A. Evans. 1996. An In­or­di­nate Fond­ness for Bee­tles. New York: Henry Holt and Com­pany, Inc..

Bor­ror, D., D. Long, C. Triple­horn. 1981. An In­tro­duc­tion to the Study of In­sects. Philadel­phia: Saun­ders Col­lege Pub­lish­ing.

Bor­ror, D., R. White. 1970. A Field Guide to In­sects. Boston: Houghton Mif­flin Com­pany.

Buchs­baum, R., M. Buchs­baum, J. Pearse, V. Pearse. 1987. An­i­mals With­out Back­bones. Chicago: The Uni­ver­sity of Chicago Press.

Bur­ton, J. 1968. The Ox­ford Book of In­sects. Ox­ford: Ox­ford Uni­ver­sity Press.

Essig, E. 1926. In­sects of North Amer­ica. New York: The Macmil­lan Com­pany.

Milne, L., M. Milne. 1980. The Au­dobon So­ci­ety Field Guide to NOrth Amer­icna In­sects and Spi­ders. New York: Al­fred A. Knopf.

Wald­bauer, G. 2000. Mil­lions of Mon­archs, Bunches of Bee­tles. Cam­bridge, Mass­a­chu­setts: Har­vard Uni­ver­sity Press.

Wee­den, , Shel­ton, Li, Hoff­man. 2004. "Cor­nell Uni­ver­sity, Bi­o­log­i­cal Con­trol" (On-line). Ac­cessed April 07, 2005 at http://​www.​nysaes.​cornell.​edu/​ent/​biocontrol/​predators/​rodolia_​cardinalis.​html.