Geographic Range
Eudyptula minor
is found throughout the southern coast of Australia and as far north as the South
Solitary Island off the coast of New South Wales. They are also native to the coasts
of New Zealand.
Eudyptula minor
has six recognized subspecies.
E. m. novaehollandia
is geographically located in Australia. The other five subspecies,
E. m. iredaei
,
E. m. variabilis
,
E. m. albosignata
,
E. m. minor
,
E. m. chathamensis
, are distributed around the country of New Zealand.
- Biogeographic Regions
- australian
- pacific ocean
Habitat
When on land,
Eudyptula minor
inhabits coastal habitats with good nesting conditions. Little penguins nest in
burrows dug in bare sand or under vegetation. If the ground is too soft to hold a
burrow, these penguins also nest in caves and rock crevices. Habitats include rocky
coastline, savanna, scrub forest or forests. Little penguins are marine seabirds and
spend the majority of their lives swimming underwater.
- Habitat Regions
- temperate
- saltwater or marine
- Terrestrial Biomes
- chaparral
- scrub forest
- Aquatic Biomes
- coastal
- Other Habitat Features
- caves
Physical Description
As the smallest penguin in the world, this flightless bird stands at an average height of 30 cm and has a weight of 1.1 to 1.2 kg. It has a black bill with an average length of 35 mm and eyes ranging from silver to blue, grey, and hazel. Its chin and throat are white along with the underside of its flippers and torso. The top of the head, neck and dorsal side of its flippers and torso are an indigo-blue. The color of the penguin’s feathers can become duller with age, and the color of their undersides can range from white to gray to brown. Sexual dimorphism is not pronounced in this species. Males are larger and have longer and deeper bills than females. Males have an average bill length of 35.7 mm and an average bill depth of 15.4 mm. Females have an average bill length of 34.5 mm and an average depth of 14.1 mm. Flipper length is similar in both genders with an average of 117.5 mm.
Juveniles have a dorsal plumage that is a brighter light blue than the indigo-blue
of the adults. The juveniles also have thinner and shorter beaks.
- Other Physical Features
- endothermic
- homoiothermic
- bilateral symmetry
- Sexual Dimorphism
- sexes alike
- male larger
Reproduction
Courtship begins with male little penguins performing courtship displays and giving mating calls. A male will hold his body in an upright position with flippers above his back, neck stretched, and head upright facing the sky. The male then emits a braying sound. These displays may be performed alone or in a group of unmated males. Occasionally the male will perform in front of a nest he constructed. After a female chooses a male, they perform a display together. One individual stands upright and spreads its flippers with head bowed, which signals the other bird to follow and they walk in small circles around the nest, braying as they go. After this display by male and female, copulation takes place.
Little penguins form monogamous pairs and retention of mated pairs from year to year
is high in this species. Pairs are likely to split up only after an unsuccessful nesting
attempt or death.
- Mating System
- monogamous
Little penguins breed from June to October in loose colonies. They may nest in ground
burrows, rocky cliffs or caves, where they lay a clutch of 1 to 2 eggs. The eggs are
smooth and white in appearance. They have an average weight of 53 g and an average
diameter of 42.0 mm. Incubation occurs for 31 to 40 days and the newly hatched chicks
are an average weight of 36 to 47 g. The chicks are semi-altricial thus are born with
downy feathers, require brooding, are unable to leave the nest, and are unable to
feed themselves. After the young hatch, the next 18 to 38 days are termed the "guard
period" for penguins during which time both parents brood the young, trading off every
3 to 4 days. After the initial guard period, the parents relax their duties and guard
chicks only at night. Fledging occurs when the chick is 50 to 65 days old and at this
time it has grown to between 800 g to 1150 g. Juveniles reach full independence at
57 to 78 days old. Most juvenile penguins reach reproductive maturity at 3 years old.
The breeding cycle of
Eudyptula minor
is variable depending on nesting location and many other environmental factors. Nutrition,
age, breeding date can influence the timing of the breeding cycle and nesting success.
A lack of nutrition has been shown to delay the breeding process. The probability
of breeding success also increases with age. This trend is attributed to the fact
that older penguins have more experience, which increases the chance of fledgling
survival. Little penguins can lay multiple clutches if the first clutch was a failure
or if the adults raised their first fledglings early in the breeding season.
- Key Reproductive Features
- iteroparous
- seasonal breeding
- gonochoric/gonochoristic/dioecious (sexes separate)
- sexual
- oviparous
Both sexes take responsibilities in the breeding process. Both male and female penguins
may build their nest together, but the male may have a greater role in physically
building the burrow. The female often takes on a larger role in the incubation stage,
but the male still helps by exchanging duties with the female every 3 to 4 days. After
chicks are born, both parents continue to brood the young during the "guard period."
Again, parents swap guarding duties every 3 to 4 days so that one broods the chicks
while the other forages. After several weeks, parents decrease guarding time to only
at night. Chicks fledge after 50 to 65 days at which time they leave the nest and
do not return for several days. Juveniles reach independence from their parents at
57 to 78 days old.
- Parental Investment
- altricial
- male parental care
- female parental care
-
pre-fertilization
- provisioning
-
protecting
- female
-
pre-hatching/birth
-
provisioning
- male
- female
-
protecting
- male
- female
-
provisioning
-
pre-weaning/fledging
-
provisioning
- male
- female
-
protecting
- male
- female
-
provisioning
-
pre-independence
-
provisioning
- male
- female
-
protecting
- male
- female
-
provisioning
Lifespan/Longevity
Little penguins live an average of 6 years. However a banded little penguin has been
recaptured the age of 25 years and 8 months old. Data on the lifespan of the bird
in captivity could not be found.
Behavior
Eudyptula minor has over a dozen different aggressive behavior displays. These displays can ultimately be divided into 4 different categories including stationary warning displays, rapidly advancing towards the intruder, brief physical contact and physical attacks. All four behaviors include a different type of physical display and vocalization.
The stationary warning displays occurs when the threat is 1 to 3 m away from the penguin.
The penguin spreads its flippers, holding its body erect and giving the intruder a
direct look accompanied by a loud vocalization. When the penguin rapidly advances
towards the intruder, it walks quickly or lunges towards the intruder with a bray-like
call. Brief physical contact can range from touching bills to slapping the intruder
with a flipper. If the penguin is in its burrow, it lunges out to peck the intruder
with its bill. If the intruder does not retreat, penguins resort to physical attacks
that include biting and beating with flippers.
Little penguins are considered the most nocturnal penguins, but generally spend all
day foraging at sea and return to land to roost at dusk. In the breeding season, penguins
swim out only an average of 8 to 9 km from shore for about 12 to 18 hours at a time.
These short trips are probably because chicks have a limited ability to thermoregulate
and need to be fed constantly. During the non-breeding season, penguins can take long
distance trips of up to 710 km, but in most cases they continue to stay within 20
km of shore. Little penguins have to use a greater amount of energy to dive into the
water than larger penguins, and although they can dive up to 67 m in depth, they mostly
remain within 5 m of the surface. When the penguins return to shore from the sea,
they parade back to their nests in groups. In good breeding years, these penguins
form these groups in a nonrandom fashion, seeking out the same individuals they paraded
with in the morning to parade back with at dusk.
- Key Behaviors
- fossorial
- natatorial
- nocturnal
- motile
- sedentary
- territorial
- colonial
Home Range
Burrow nests are usually over 2 m apart in little penguin colonies. However, when
the penguins nest in caves the nests are often closer than 2 m apart.
Communication and Perception
Eudyptula minor is a nocturnal species and is highly vocal during the night while roosting. The sound of their calls can range from a low rumble to a trumpet-like noise. Their song can be used for several functions, including attracting mates. Each little penguin has a distinctive individual song that is used by parents and siblings to distinguish one another from strangers. Calls can also be used with an aggressive intent against an intruder around a penguin's nest.
Little penguins perform unique courtship displays. Males take a particular stance, with heads facing up and wings back, while braying to females. If the female accepts, she will join the male in a courtship "dance" where they march in circles together and make braying calls.
Like all birds, little penguins perceive their environments through visual, auditory,
tactile and chemical stimuli.
- Other Communication Modes
- duets
Food Habits
Eudyptula minor
is mainly piscivorous and employs a pursuit-diving technique to catch prey in shallow
depths. The majority of its diet is composed of
Clupeiformes
fish, such as anchovies and sardines. The variety of fish consumed depends on the
locality of the penguin. This species also preys on small squid, octopi and crustaceans.
It has been observed that in recent years the number of prey available is decreasing.
This results in longer foraging trips for the penguin, greater energy expenditures,
and can ultimately decrease population sizes.
- Animal Foods
- fish
- mollusks
- aquatic crustaceans
Predation
Key predators of little penguins are introduced species. These include
dogs
,
weasels
,
rats
,
foxes
and
cats
.
Pacific gulls
and
King's skinks
are natural predators that eat the eggs and young of little penguins. In an effort
to decrease predation, little penguins move in groups to and from the ocean. This
anti-predator technique occurs a few hours before dawn and a few hours after dusk
when it is dark. As penguins are less mobile on land, making mass land movements
under the cover of darkness is likely another method used to avoid predation. Despite
these techniques, adult little penguins often fall prey to sharks, seals, and
orca whales
.
Ecosystem Roles
Eudyptula minor
plays multiple roles in its ecosystem as a predator and a host to parasites. It preys
on small fish, squids, octopi, or occasionally crustaceans and likely impacts these
populations. Little penguin eggs and chicks are food sources to local populations
of dogs, rats, cats, and other introduced predators. Adult little penguins fall prey
to sharks, seals, and orca whales and are a valuable food source to these predators.
In recent years, a new species of feather mite,
Ingrassia eudyptula
, has been discovered which is believed to parasitize
Eudyptula minor
. These mites eat preening oil on the feathers of the penguin.
- Feather mites ( Ingrassia eudyptula )
Economic Importance for Humans: Positive
The penguin parade of the
Eudyptula minor
is a popular tourist attraction. It has been recorded that 500,000 tourists annually
come to watch the colony of penguins parade to and from the water at Phillip Island.
This specific species is also of great interest to scientists because of their small
sizes and the increased amounts of energy needed to survive, especially in cold temperatures.
This subject is important in the study of thermoregulation in endotherms, and the
penguin's physical characteristics allow scientists to use this penguin in comparisons
with other endotherms.
- Positive Impacts
- ecotourism
- research and education
Economic Importance for Humans: Negative
There are no known effects of little penguins on humans.
Conservation Status
Currently
Eudyptula minor
is not considered to be threatened by extinction. It is believed that the global
population of these birds averages around 1,000,000 individuals. Their population
is declining, however, due to introduced predators, decreasing populations of prey
and oil spills. The intensity of industrial fisheries results in low prey densities
for penguins and other piscivores. Factors such as human settlement, coastal erosion,
and pollution have also affected the breeding habitats of these birds.
The subspecies
E. m. albosignata
is now considered endangered. It is only found on the Banks Peninsula on South Island,
New Zealand.
Additional Links
Contributors
Ashley Chung (author), University of Michigan-Ann Arbor, Phil Myers (editor), University of Michigan-Ann Arbor, Rachelle Sterling (editor), Special Projects.
- Australian
-
Living in Australia, New Zealand, Tasmania, New Guinea and associated islands.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- Pacific Ocean
-
body of water between the southern ocean (above 60 degrees south latitude), Australia, Asia, and the western hemisphere. This is the world's largest ocean, covering about 28% of the world's surface.
- native range
-
the area in which the animal is naturally found, the region in which it is endemic.
- temperate
-
that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).
- saltwater or marine
-
mainly lives in oceans, seas, or other bodies of salt water.
- chaparral
-
Found in coastal areas between 30 and 40 degrees latitude, in areas with a Mediterranean climate. Vegetation is dominated by stands of dense, spiny shrubs with tough (hard or waxy) evergreen leaves. May be maintained by periodic fire. In South America it includes the scrub ecotone between forest and paramo.
- scrub forest
-
scrub forests develop in areas that experience dry seasons.
- coastal
-
the nearshore aquatic habitats near a coast, or shoreline.
- endothermic
-
animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.
- bilateral symmetry
-
having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.
- monogamous
-
Having one mate at a time.
- iteroparous
-
offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).
- seasonal breeding
-
breeding is confined to a particular season
- sexual
-
reproduction that includes combining the genetic contribution of two individuals, a male and a female
- oviparous
-
reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.
- altricial
-
young are born in a relatively underdeveloped state; they are unable to feed or care for themselves or locomote independently for a period of time after birth/hatching. In birds, naked and helpless after hatching.
- male parental care
-
parental care is carried out by males
- female parental care
-
parental care is carried out by females
- fossorial
-
Referring to a burrowing life-style or behavior, specialized for digging or burrowing.
- natatorial
-
specialized for swimming
- nocturnal
-
active during the night
- motile
-
having the capacity to move from one place to another.
- sedentary
-
remains in the same area
- territorial
-
defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement
- colonial
-
used loosely to describe any group of organisms living together or in close proximity to each other - for example nesting shorebirds that live in large colonies. More specifically refers to a group of organisms in which members act as specialized subunits (a continuous, modular society) - as in clonal organisms.
- visual
-
uses sight to communicate
- acoustic
-
uses sound to communicate
- duets
-
to jointly display, usually with sounds in a highly coordinated fashion, at the same time as one other individual of the same species, often a mate
- ecotourism
-
humans benefit economically by promoting tourism that focuses on the appreciation of natural areas or animals. Ecotourism implies that there are existing programs that profit from the appreciation of natural areas or animals.
- carnivore
-
an animal that mainly eats meat
- piscivore
-
an animal that mainly eats fish
- visual
-
uses sight to communicate
- tactile
-
uses touch to communicate
- acoustic
-
uses sound to communicate
- chemical
-
uses smells or other chemicals to communicate
References
Daniel, T., A. Chiaradia, M. Logan, G. Quinn, R. Reina. 2007. Synchronized group association in little penguins, Eudyptula minor . Animal Behaviour , 74: 1241-1248.
Davis, L., M. Renner. 2003. Penguins . New Haven and London: Yale University Press.
Fallow, P., A. Chiaradia, Y. Ropert-Coudert, A. Kato, R. Reina. 2009. Flipper Bands Modify the Short-Term Diving Behavior of Little Penguins. Journal of Wildlife Management , 73/8: 1348-1354.
Heber, S., K. Wilson, L. Molles. 2008. Breeding biology and breeding success of blue penguin ( Eudyptula minor ) on the West Coat of New Zealand's South Island. New Zealand Journal of Zoology , 35: 63-71.
Hoskins, A., P. Dann, Y. Ropert-Coudert, A. Kato, A. Chiaradia, D. Costa, J. Arnould. 2008. Foraging behaviour and habitat selection of the little penguin Eudyptula minor during early chick rearing in Bass Strait, Australia. Marine Ecology-Progress Series , 366: 293-303.
Jouventin, P., T. Aubin. 2000. Acoustic convergence between two nocturnal burrowing seabirds: experiments with a penguin Eudyptula minor and a shearwater Puffinus tenuirostris . Ibis , 142: 645-656.
Kato, A., Y. Ropert-Coudert, A. Chiaradia. 2008. Regulation of Trip Duration by an Inshore Forager, the Little Penguin ( Eudyptula Minor ), During Incubation. The Auk , 125/3: 588-593.
Knight, C., T. Rogers. 2004. Factors influencing fledgling production in little penguins. Wildlife Research , 31: 339-344.
Mironov, S., H. Proctor. 2008. The Probable Association of Feather Mites of the Genus Ingrassia (Analgoidea: Xolalgidae) with the Blue Penguin Eudyptula minor (Aves: Sphenisciformes) in Australia. The Journal of Parasitology , 94/6: 1243-1248.
Miyazaki, M., J. Waas. 2002. 'Last Word' Effects of Male Advertising Calls on Female Preference in Little Blue Penguins. Behaviour , 139/11-12: 1413-1423.
Nakagawa, S., J. Waas, M. Miyazaki. 2001. Heart rate changes reveal that little blue penguin chicks ( Eudyptula minor ) can use vocal signatures to discriminate familiar from unfamiliar chicks. Behavioral Ecology and Sociobiology , 50: 180-188.
Nisbet, I., I. Dann. 2009. Reproductive performance of little penguins Eudyptula minor in relation to year, age, pair-bond duration, breeding date and individual quality. Journal of Avian Biology , 40/3: 296-308. Accessed February 22, 2010 at http://dx.doi.org/10.1111/j.1600-048X.2008.04563.x .
Overeem, R., A. Peucker, C. Austin, P. Dann, C. Burridge. 2008. Contrasting genetic structuring between colonies of the World's smallest penguin, Eudyptula minor . Conservation Genetics , 9/4: 893-905.
Peucker, A., P. Dann, C. Burridge. 2009. Range-Wide Phylogeography of the Little Penguin ( Eudyptula minor ): Evidence of Long-Distance Dispersal. The Auk , 126/2: 397-408.
Ropert-Coudert, Y., A. Kato, A. Chiaradia. 2009. Impact of small-scale environmental perturbations on local marine food resources: a case study of a predator, the little penguin. Proceedings of The Royal Society , 276: 4105-4109.
Sergent, N., T. Rogers, M. Cunninghman. 2004. Influence of biological and ecological factors on hematological values in wild Little Penguins, Eudyptula minor . Comparative Biochemistry and Physiology - Part A: Molecular & Integrative Physiology , 138/3: 333-339.
Thomas, ., R. Fordyce. 2007. The Heterothermic Loophole Exploited by Penguins. Australian Journal of Zoology , 55/5: 317-321.
Williams, T. 1995. The Penguins: Spheniscidae . New York: Oxford University Press.
2009. "Little Penguin, Eudyptula minor " (On-line). Parks & Wildlife Service, Tasmania. Accessed March 19, 2010 at http://www.parks.tas.gov.au/index.aspx?base=5091 .